The term represents prey reproduction and limits growth by available food, giving carrying capacity in the absence of hunters. The loss represents encounters between prey and hunters. For hunters, is prey-supported growth and is mortality; their per-capita growth changes sign at prey population . The coordinate axes are invariant, and positive initial populations remain positive.
For , the physical equilibrium points are the origin, , and when . The Jacobian matrix is
At its eigenvalues are and , so the origin is always a saddle equilibrium, attracting along the hunter-only axis and repelling along the prey-only axis. At the eigenvalues are
It is a saddle equilibrium for and an asymptotically stable node for .
For , the coexistence Jacobian matrix is
Its trace is negative and determinant positive. Its discriminant is negative, so is an asymptotically stable focus. Interior trajectories approach it in damped, counterclockwise oscillations in the plane. Both populations ultimately coexist.
To justify the global interior fate for this logistic predator-prey model, let and use the Lyapunov function
It is nonnegative and its sublevel sets are compact inside the positive quadrant. Differentiation gives the explicit cancellation
The only invariant subset on which this vanishes is : remaining at requires . LaSalle's invariance principle therefore proves that every strictly positive initial state converges to this coexistence point.
For , the formal coexistence point has negative and is outside the physical state space. The only physical equilibria are the saddle equilibrium at the origin and the stable prey-only node. Since , positive prey has . Eventually with a fixed margin, so hunter population decays exponentially. When is sufficiently small, comparison with gives for every . Thus and for positive initial prey: hunters die out and prey reaches carrying capacity.
On the boundary, hunter-only initial data approach the origin; prey-only positive data approach in either parameter range. The sketches show representative values in the two requested ranges. Dashed and dotted lines are the prey and hunter nullclines; filled dots are attractors and open dots are saddle equilibria.
Figure 1.
Predator-prey phase portraits showing stable coexistence for b=0.25 and hunter extinction for b=1.5
. Details.