For a straight, unloaded inextensible filament with small transverse displacement, resistive-force theory gives viscous force density . Variation of the bending energy gives , while the induced filament tension is higher order. Their instantaneous balance yields . It is an overdamped fourth-order diffusion equation: short-wavelength bends relax much faster than long ones. Wiggins and Goldstein's flexive-propulsion paper develops the elastic-wave mechanism for low-Reynolds-number propulsion.
Let positive transverse displacement point along a uniform load per unit length . The filament bending modulus gives energy . The variational derivative is , so resistive-force theory gives . Clamping imposes , while a free tip imposes . The steady solution is
Its tip displacement is . For a submerged cylindrical filament under Newtonian gravity, buoyancy gives . This linear approximation requires small slope and negligible inertia.
For a uniform-load bending of a clamped filament, total load and tip displacement obey Hooke's law in the form , with . This stiffness depends on the specified distributed loading. A concentrated tip force instead gives , so equal total forces with different distributions are not interchangeable.
The anisotropic-drag contribution to axial force from a small-slope transverse motion is . Using the small-slope elastohydrodynamic filament equation and integration by parts gives
Thus this propulsive contribution is determined by endpoint slope, bending moment, and shear. A separately imposed axial translation adds ordinary longitudinal drag. Exact second-order longitudinal motion required by inextensibility also contributes to instantaneous total drag, but its time-derivative contribution averages to zero over a deformation cycle.
In units of elastohydrodynamic penetration length and inverse forcing frequency, . Prescribing , zero bending moment , and decay at infinity gives
The two components have phase velocities and . Their unequal spatial attenuation produces a nonreciprocal shape cycle even though the endpoint actuation is reciprocal.

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