The first equation is mass conservation for an incompressible flow whose area may change: local area storage balances the divergence of the volume flux . The two left-hand momentum terms are local acceleration and convective acceleration, while is the driving pressure gradient and models viscous drag per unit mass. The pressure-area law of a collapsible tube expresses transverse wall balance using internal minus prescribed external pressure.
For example, choose a reference area , length and velocity . If dimensional variables are , takeThe dimensionless external pressure uses the same scale. A dimensional friction rate becomes , and the dimensional wall pressure law is divided by . These choices make both inertia and the pressure-gradient coefficient unity; other equivalent reference scales are possible. On the mechanically stable wall branch, increasing area requires increased transmural pressure, whereas an enlarged tube reduces resistance. Thus .
Steady mass conservation makes . For uniform area, is constant and the pressure-area law of a collapsible tube gives . The acceleration vanishes, so momentum balance requiresThe external pressure must therefore decrease linearly along a positive-flow tube. Its additive constant is free; the constant transmural pressure is .
Put , , and . Let the area and velocity perturbations have amplitudes and . Linearizing mass conservation and momentum, and using , givesFor nonzero , eliminate to obtain the friction-induced instability of a collapsible-tube flow relationAt the stated threshold , it factors asTherefore one mode has real , while the other has and decays. The PDF's real-frequency statement is valid for the neutral branch, not for both roots. The Fourier convention makes positive imaginary frequency mean growth.
Complete the square in the dispersion relation:If the imaginary part of the square root has magnitude , thenOne root grows exactly when . For , squaring the equivalent positive inequality gives . Thus positive flow is unstable whenever . This proves the entire stated range, not just a small neighborhood of threshold. Zero wavenumber has only a neutral uniform-area change and a damped velocity mode.
Near threshold write and follow the neutral branch as . Substitution yieldsTaking its imaginary part gives the growth rateArea increases reduce friction, causing a velocity increase that feeds the area disturbance through mass conservation. The pressure-driven base flow supplies the energy; the positive friction coefficient does not by itself prevent instability of that maintained flow.
In the travelling coordinate , mass conservation becomes . Thus the conserved relative flux is andFor a periodic peristaltic pumping solution, the neglected-inertia momentum equation isTake to be the spatial mean area, so . This normalization, and zero net pressure drop over a period, are implicit in the specified periodic forcing. The constant leading state has no pressure gradient, giving . Put and . At first order,Period averaging gives , and the zero-mean solution isFor the second-order mean balance, expand the velocity and friction:Averaging the exact pressure derivative gives zero, so . Hence the weak peristaltic pumping of a collapsible tube result isThe mean laboratory volumetric flow rate is , forward with the wave when . The negative leading relative flux is therefore consistent with positive mean pumping.
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