For a Laplacian mode with , the linearized chemotaxis system isIts trace is negative for all . Writing , its determinant isInstability occurs when this determinant is negative, because then one real eigenvalue is positive. A quadratic with positive leading and constant coefficients is negative for some precisely when its middle coefficient is negative and its discriminant positive. These conditions combine toThis proves the printed threshold. In a bounded spatial domain, only its permitted Laplacian eigenmodes are available; the continuous- threshold describes the onset when wavenumbers can vary freely.
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