Let be arclength, the unit tangent, and the local filament velocity relative to the fluid. The resistive-force theory force per unit arclength, exerted by the fluid on the filament, is
The parallel and perpendicular drag coefficients of a slender filament are proportional to dynamic viscosity. For radius and a relevant axial cutoff length , the anisotropic slender-filament drag from Stokeslet integration gives the leading logarithmic forms
The cutoff may be set by filament length or deformation wavelength. End shape and the choice of cutoff change order-one constants inside the logarithms. Slender-body theory also includes nonlocal interactions that the local resistive-force theory approximation omits. Typically , with ratio tending to two in the slender limit; this drag anisotropy enables propulsion.
Write . For a small-amplitude planar deformation, and the leading transverse velocity is . Exact inextensibility requires the quadratic longitudinal displacement of an inextensible planar filament: if and , then to this order. Its velocity is quadratic and has zero time average for a periodic material cycle without mean axial drift. Thus the fixed material abscissa in the question is leading-order notation; this correction does not change the averaged propulsive term below. An imposed mean translation would contribute its separate ordinary axial drag.
To leading nonzero order,
The time-averaged deformation-induced propulsive force of a periodic planar filament and power of a periodic planar filament are consequently
These are the leading quadratic functionals. Their signs refer to force on the filament. For example, gives negative , opposing the direction of wave propagation.
Let be an arbitrary smooth variation periodic in both variables. The first variation of the constrained functional is
Using integration by parts in and , every boundary term cancels by periodic boundary conditions. Therefore
The Euler-Lagrange equation is
For a nontrivial stroke with nonzero thrust, . Set and . Then , whose method of characteristics solution is . Integrating in time gives the travelling-wave stationarity of planar filament propulsion:
Here is an arbitrary static periodic component. It contributes neither power nor mean propulsion because the time average of vanishes. It can be set to zero when describing the active stroke. The wave must be compatible with both specified periods: .
For a pure travelling wave, its leading force and power are
Thus positive propulsion uses a wave travelling in the negative direction. Any smooth interior maximizer satisfying only the stated power constraint must have this travelling active deformation. If the drag were isotropic, and there would be no leading propulsion.
There is a qualification to the printed maximization claim: the variation proves stationarity, not the existence of an unrestricted global maximum. The bandwidth limitation in fixed-power filament optimization can be seen directly. Set , , and consider . The leading expressions give
At fixed power the formal quadratic objective increases with . Eventually that sequence violates the small-slope approximation, but the approximation supplies no numerical slope or wavelength cutoff with which to define the global optimization. Even within a strict small-slope neighbourhood, a sufficiently small admixture of a higher spatial harmonic can improve a candidate interior maximum while keeping power fixed. For example, retains the fundamental periods and exactly the same leading power, while its force is multiplied by . For fixed , taking and small keeps every slope small.
With an explicit admissible Fourier bandwidth or an additional geometric constraint, the intended conclusion can be made precise. For a finite set of nonstatic Fourier series modes proportional to , with wavenumbers and frequencies ,
This is a weighted average of . A largest allowed ratio is attained by a single travelling mode, or by modes sharing the same phase speed; their superposition is still a travelling wave. Thus the Euler-Lagrange calculation yields the requested travelling-wave form, while a global maximum requires a specified admissible shape class.

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