Write the membrane height above its mean plane as a sum of Fourier modes, . The supplied average is a thermal mean-squared modal height; for complex Fourier coefficients it is understood as , with for a real height field. The terms are:
- is the thermal energy, with Boltzmann constant and absolute temperature .
- is the projected membrane area used in the Fourier normalization.
- is the in-plane wavenumber; the corresponding wavelength is .
- is the membrane tension, in .
- is the membrane bending modulus, in joules.
In the small-slope approximation, Helfrich membrane energy containsThe equipartition theorem, with the independent real modes counted consistently, gives the stated membrane fluctuation spectrum under tension. Its units are length squared: and both have units , so has units .
At small length scales, is large and the bending term dominates:The fluctuation amplitude is controlled mainly by the membrane bending modulus and thermal energy, rather than tension. The crossover occurs when , henceThe bending-dominated regime extends up to the membrane elastic length when scale means , or up to when it means wavelength. At still larger scales tension gives a spectrum. At the opposite end, the continuum formula requires wavelengths larger than the microscopic membrane thickness or molecular cutoff. Without numerical and , no numerical crossover scale is determined.
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