For a nearly flat membrane represented by height , the quadratic bending-and-tension energy per invariant transverse length is
Its free equilibrium equation is .
For a membrane of bending modulus adhering with energy per area to a cylinder of radius , compares adhesion with bending cost.
A membrane-mediated interaction potential is the separation-dependent energy produced when deformation fields around two membrane-bound objects overlap. Its sign and range depend on membrane tension, bending stiffness, inclusion geometry, and which side of the membrane each object occupies.

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