Stretch and twist coefficients of an elastic filament ($K_b=k_{B}TA,S=k_{B}TB,K_t=k_{B}TC$)

For an elastic filament with reference arclength $s$, curvature $c$, axial strain $ϵ$ and excess twist rate $\Omega$, a harmonic energy density is $(K_b{c}^2+S{ϵ}^2+K_t{\Omega }^2)/2$. Bending and torsional rigidities have units of energy times length; the stretch modulus has units of force. Factoring out $k_{B}T$ therefore makes $A,C$ lengths but $B$ an inverse length. Axial strain under tension is approximately $F/S$. Twist is controlled by torque and rotational boundary conditions, not force alone. A three-dimensional thermally fluctuating worm-like chain has persistence length $K_b/(k_{B}T)$.