The per-capita net growth rate is . Thus is the low-density per-capita birth or recruitment rate, is the constant per-capita death or removal rate, and controls density-dependent suppression of recruitment. Both and have units of inverse time; has units of inverse population squared. The population scale halves the recruitment rate from its low-density value. Larger makes suppression important at smaller population sizes. These are interpretations of this phenomenological population dynamics model, rather than independently specified biological mechanisms.

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