For , . When , the roots of are real and lie strictly between zero and one. When , they are a complex-conjugate pair of modulus . Thus the positive steady state is linearly asymptotically stable exactly when , and unstable when . Since increases strictly with , the crossing occurs atAt the crossing,Every nonzero real linearized perturbation has period six. This establishes the intended linear stability threshold and linearized period six.
The nonlinear period-six claim in the printed paper does not follow, and is false as a local bifurcating-branch claim for this map. One can check this directly rather than infer a nonlinear period from the eigenvalues. At normalize , . The exact planar map isIts cubic Taylor expansion isHere is a concrete check of the sixfold return. Put , , andComposing the displayed cubic expansion six times cancels all quadratic terms and givesIn particularThe quartic coefficient is strictly negative. Hence, sufficiently near the equilibrium, every nonzero point has smaller after six iterations: the critical nonlinear map has no nearby nontrivial period-six orbit, and the equilibrium itself is locally attracting despite its neutral linearization.
One can also rule out a branch of small period-six orbits on either side of the crossing. Normalize about the positive equilibrium at each nearby , and put . The derivative of the sixth power of at is . Smooth dependence on therefore givesThe leading matrix has determinant . Both terms could vanish with only if , butThus its norm on nonzero is bounded below by a positive constant times , and the smaller remainder cannot cancel it. There is no nearby nonzero solution of . The eigenvalues satisfy the nonresonance conditions through order four, their moduli cross outward with increasing , and the cubic radial coefficient is negative. The standard Neimark–Sacker bifurcation theorem therefore gives a supercritical invariant closed curve. Its rotation is near the linear resonance ; it is not a local branch of exact period-six orbits.
Finally, the discrete population map does not preserve nonnegativity for arbitrary large lagged populations: can be negative. The positive-equilibrium stability conclusions above are local and do not assume a global positivity property that the formula lacks.
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