In coordinate order the Jacobian matrix isAt the origin it is , giving a repelling node equilibrium; when the repeated positive eigenvalue is semisimple. At coexistence it isIts trace is , determinant , and characteristic discriminant . Hence both eigenvalues are real and negative, proving a stable node equilibrium for every .
The other equilibria are and . Their triangular Jacobians have eigenvalues and respectively. Thus both single-species equilibria are saddles, with their attracting directions on the corresponding coordinate axes and unstable directions allowing invasion by the other species. There are no further nonnegative equilibria because the two non-axis nullclines intersect only at .
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