For , has eigenvectors and with eigenvalues and . The Neumann spatial modes are , . Combining each with those two reaction directions gives temporal ratesEvery linear perturbation therefore decays at least as fast as . The slowest mode is the spatially uniform antisymmetric change of the two biological populations. ThusSmall nonlinear perturbations have the same bound, not just a formal mode calculation. In one spatial dimension take sufficiently small smooth Neumann data, or small data in a norm such as that controls quadratic products. The linear semigroup decays at rate , and the nonlinear remainder is quadratic. Standard small-data variation-of-constants estimates first give decay for any fixed . Choose ; then the remainder is , and its Duhamel convolution with is because . This sharpens the preliminary bound to the displayed rate. Special initial data with no slow-mode component may decay faster, but the general estimate cannot be improved uniformly.
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