The coexistence Jacobian matrix calculated in part (a) has negative diagonal entries and positive determinant. For any spatial mode with squared wave number , the linearized matrix is . Its trace is more negative than before, and its determinant isHence both eigenvalues retain negative real parts for every mode and every . This is the mechanism by which negative self-response excludes two-species Turing instability, excluding diffusion-driven instability using only the reaction Jacobian matrix already computed, together with the diagonal diffusive damping. In particular unequal diffusion is not enough to produce a Turing pattern when both local self-responses are negative in this two-species competitive model.
For , has eigenvectors and with eigenvalues and . The Neumann spatial modes are , . Combining each with those two reaction directions gives temporal ratesEvery linear perturbation therefore decays at least as fast as . The slowest mode is the spatially uniform antisymmetric change of the two biological populations. ThusSmall nonlinear perturbations have the same bound, not just a formal mode calculation. In one spatial dimension take sufficiently small smooth Neumann data, or small data in a norm such as that controls quadratic products. The linear semigroup decays at rate , and the nonlinear remainder is quadratic. Standard small-data variation-of-constants estimates first give decay for any fixed . Choose ; then the remainder is , and its Duhamel convolution with is because . This sharpens the preliminary bound to the displayed rate. Special initial data with no slow-mode component may decay faster, but the general estimate cannot be improved uniformly.
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