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A leading short-distance Standard Model contribution to neutral D-meson mixing is a box Feynman diagram with two W bosons and two internal down-type quark lines:

The incoming $c\bar{u}$ pair becomes $u\bar{c}$; the internal labels run over the down quark, strange quark and bottom quark. Each corner is a weak charged current vertex. Four such vertices make the contribution of order $g^4$. Summing the internal flavours produces Cabibbo-Kobayashi-Maskawa matrix factors and the Glashow-Iliopoulos-Maiani mechanism: the flavour-independent term cancels by ${\sum }_i{V}_{ci}^{\ast }{V}_{ui}=0$. This box Feynman diagram is one contribution; long-distance intermediate hadron states can also contribute to neutral D-meson mixing.

The antiparticle of the strange quark has electric charge $+1/3$ and baryon number $-1/3$.