Carrying capacity 2026-10-14
A carrying capacity is an abundance scale at which a specified environment's net population growth vanishes. In the logistic growth equation, positive unharvested populations approach . It depends on the species, resources and environment rather than being an immutable universal population limit.
The useful distinction is between a growth rate and a sustainable abundance. In a logistic differential equation, controls how quickly a population responds, while the carrying capacity sets the positive equilibrium point. Increasing with fixed accelerates approach without raising that equilibrium. Increasing changes the destination. The logistic differential equation diagrams show populations approaching the same carrying capacity from below and above.
In ecological use, the carrying capacity describes what a particular environment can support under specified conditions. Nutrient supply, available space, and accumulated waste can change those conditions; it is not a permanent property of a species alone. A population temporarily above a carrying capacity is possible. In the simplest constant- logistic growth equation, its net growth becomes negative; real population dynamics can include delays, seasonal variation, and changing resources that the basic mathematical model omits. [ref].
For , the positive amplitude ordinary differential equation has solution . This follows from the logistic differential equation for . Every approaches as a monotone function, and the positive distance to this limit decreases; an oscillating complex wave carrying this amplitude has no ordinary monotone ordering.
The Forward Euler method applied to the logistic differential equation becomes the logistic map after , . The positive equilibrium point of the differential equation has perturbations decaying as . Its discrete fixed-point multiplier is : for , the discretization has an artificial oscillatory instability, leading initially to alternation on successive steps. This contrasts with the positive perturbation multiplier of the exact time-step map.
A growing linear mode can make a formally small derivative nonlinearity order one at large distance. After the crossover scaling, the equation reduces to a logistic differential equation in for . Matching at small gives and the displayed primitive with . It changes from quadratic growth to linear growth without a finite turning singularity.
Logistic differential equation Created 2026-09-28 Updated 2026-10-14
The logistic differential equation is the nonlinear ordinary differential equation
As a mathematical model of population dynamics, it treats abundance as a continuous nonnegative quantity. The constant has units of inverse time, while the carrying capacity has the same units as . The useful idea is to combine growth resembling exponential growth at low abundance with a slowing approach to a finite abundance. The three trajectories below already show an important distinction: starting above the carrying capacity gives a decreasing curve, whereas starting below it gives an increasing one.
Figure 1.
Logistic trajectories approach the carrying capacity from below or above
. Here and per day. The initial conditions are ; dashed horizontal lines show the two equilibrium points and . Only the trajectory has a future inflection point, marked at and days. The other marked values are and for that same trajectory. Details.
For , dividing the derivative by abundance gives the instantaneous per-capita rate:
It decreases linearly with : it approaches at low abundance, is at , vanishes at the carrying capacity, and is negative above it. This per-capita rate differs from the total rate , which also counts how many individuals contribute. It is an instantaneous rate, rather than a fixed percentage increase over a whole day. The linear per-capita assumption and the resulting qualitative picture are developed in MIT 18.03SC's logistic-model notes.
Writing , the equilibrium points satisfy , so the constant solutions of a differential equation are and . The phase line points upward for and downward for . Also and : the criterion for fixed point stability for an autonomous differential equation makes zero an unstable equilibrium and an asymptotically stable equilibrium point. Every positive initial condition approaches as ; stays exactly zero. Instability of zero means that a small positive population grows away from it, not that a zero population spontaneously appears.
For every , the unique solution of a differential equation for all is
The denominator is positive on this domain, including when . For , this is a translated and scaled logistic function; its extension to all real times is a sigmoid function. For the coefficient of the exponential function is negative, and the future curve instead decreases toward . No nonconstant positive solution of a differential equation crosses or reaches it at finite time. A short derivation uses the dimensionless variable : the chain rule gives , so exponential decay yields . The substitution divides by , so the zero equilibrium point must be included separately.
The total growth rate is a downward-opening quadratic function of abundance. Completing the square gives its maximum without solving the differential equation:
The balance is beautiful: more individuals contribute to growth, but each contributes less, and the strongest total increase occurs halfway to the carrying capacity. This is a maximum over possible abundances. A trajectory starting above never passes through that abundance in its future.
Figure 2.
Total logistic growth is greatest at half the carrying capacity
. With and per day, the maximum is individuals per day at . The function graph crosses zero at the equilibrium points and ; beyond the rate is negative. The lower phase line has arrows toward increasing abundance below and toward decreasing abundance above , with a hollow mark for the unstable equilibrium and a filled mark for the asymptotically stable equilibrium point. Details.
The chain rule connects this maximum to the shape of the time trajectory through its second derivative:
For , the trajectory initially has convexity and then concavity, with its one future inflection point at
For , this point lies at the initial time, with concavity for . For , the inflection point belongs to the past extension, so the future trajectory has concavity throughout. Above , the decreasing trajectory has convexity and no inflection point. The two constant solutions of a differential equation have no change of concavity. Thus an entire logistic function has an S shape, but its observed future segment need not show both bends. See also [ref].
For the plotted example with , and , time measured in days gives
The initial total rate is individuals per day, and the initial per-capita rate is . The fastest total increase is individuals per day, reached at days. At that time the per-capita rate has already fallen to .
These parameters are illustrative choices, rather than an empirical fit. The equation specifies deterministic evolution under a phenomenological assumption: it summarizes how net growth depends on abundance without separately modelling every process that affects it. The carrying capacity is a parameter for a specified setting, not a universal constant for all populations. In his 1838 paper "Notice sur la loi que la population suit dans son accroissement", Correspondance mathématique et physique 10, pp. 113–121, Pierre-François Verhulst proposed , equivalent to this equation with and . He also noted that different assumed growth-retarding terms could describe the observations similarly while giving very different limiting populations. This is an early illustration of why a good-looking curve alone does not establish a unique long-term prediction.[ref]
The logistic differential equation describes continuous time. The logistic map instead advances in discrete steps and can exhibit a period-doubling cascade and Devaney chaos; the monotone positive trajectories here cannot. Their connection through the Forward Euler method is explained under Euler discretization of logistic growth, where a large step can introduce behaviour absent from the original differential equation.
Logistic growth equation 2026-10-14
The logistic growth equation applies the logistic differential equation to population dynamics. For positive growth rate and carrying capacity , an initial condition gives , where . Its per-capita growth falls linearly with abundance, and a trajectory starting below reaches an inflection point when . The exact solutions of a differential equation and their diagrams are developed under logistic differential equation. A translated and scaled logistic function has the same form. In nonlinear regression for biological size, an analogous saturation curve need not imply that circumference itself obeys a mechanistic population law.
Nonlinear ordinary differential equation Created 2026-09-25 Updated 2026-10-14
A nonlinear ordinary differential equation depends nonlinearly on the unknown function or its derivatives: it cannot be written in the form defining a linear differential equation. For example, contains a square of the unknown function, and is a logistic differential equation. A coefficient such as depending only on the independent variable does not make an ordinary differential equation nonlinear. This distinction concerns dependence on the unknown, rather than the shape of its function graph.
For a concrete initial value problem, consider , . Its solution of a differential equation is
Taking its derivative gives , and evaluating at zero checks the initial condition. The stated interval contains zero and ends at finite-time blowup: tends to positive infinity as approaches from below. Increasing the positive initial condition brings finite-time blowup closer. This is a simple mathematical example with dimensionless and , rather than a calibrated physical model.
Choose and , giving and . At they have values and , so their sum has value . On the common interval , both are valid solutions of a differential equation, but their sum fails the same ordinary differential equation:
Already at , and its derivative is , whereas would require . At , the defect is . The cross term explains the failure of the superposition principle, even though each individual solution of a differential equation is a smooth function.
Figure 1.
Two solutions of a differential equation for have a sum with a nonzero defect
. Details.
The first panel shows , and their invalid sum only up to , before either finite-time blowup. The second measures the defect in the ordinary differential equation, including the computed value at . For a homogeneous linear differential equation, the corresponding defect of a sum is zero. Nonlinear ordinary differential equations can describe state-dependent growth such as the logistic differential equation, whose negative quadratic term limits growth instead of accelerating it without bound. Recognizing the distinction tells us when we may assemble solutions of a differential equation from simpler pieces and when we must account for their interaction. See MIT's worked counterexample to nonlinear superposition.
For the logistic differential equation, separate variables to obtain away from the constant solutions. Applying the positive initial condition gives
The formula also includes the constant solution . Below positive solutions increase; above they decrease. They approach as . Equivalently, linearizing gives and . Thus zero is unstable and is asymptotically stable.
The Forward Euler method gives . Set and ; multiplying the recurrence by this scaling factor gives the logistic map
Its fixed points are and . The fixed-point multipliers are and . Since , zero is always unstable. The positive fixed point is stable for and unstable for . For , a small displacement obeys : its sign alternates and its amplitude grows, giving an oscillatory instability whose period is .
This is an artifact of Euler discretization of logistic growth. Exact small perturbations decay by the positive factor per time step; a one-dimensional autonomous differential equation cannot have a nonconstant periodic orbit, since its solutions move monotonically between equilibrium points. The artificial instability occurs when , beyond the stable step range of this discretization.
For , remove the fixed point factors from to obtain . The resulting logistic map two-cycle is
These points map to each other. Their periodic-orbit multiplier is , so the two-cycle attracts nearby iterates. The cobweb shows a small initial departure from growing with alternating sign and then settling into the finite rectangle between and . The time trace shows the corresponding finite-amplitude oscillation.
Figure 1.
Logistic-map cobweb and alternating iterates approaching the stable two-cycle at parameter 3.2
. Details.
Putting gives the local regular perturbation result . It cannot describe the whole half-line: its derivative grows like , making order one at .
The useful exact change of variables is
Direct substitution turns the equation into two autonomous differential equations,
The initial values here are . Near the origin, linearizing this system gives eigenvalues and . Equivalently, neglecting only the derivative-square nonlinearity in the original equation gives the two modes and . Their matching coefficients are
Thus the growing derivative has . Keeping these unexpanded linear-mode coefficients is convenient for matching; it does not assert that the linearized solution is the exact nonlinear one.
At the distinguished limit , set . To leading order the derivative equation becomes , the logistic differential equation. Matching as selects
Then , and the growing-mode matching gives
This proves the logistic crossover in a weakly nonlinear Euler equation. In particular the nonlinear-region leading solution is , or with the constants taken at their lowest order.
A composite asymptotic expansion retaining the small-distance stable mode and satisfying the initial value exactly is
Since , this matches when and hence at fixed as . Its derivative at one differs from the prescribed value only by , within leading-order accuracy. For it gives . Thus the composite covers the initial region, the nonlinear transition and the arbitrarily large- regime; uniform leading accuracy is understood relative to the growing solution, not as an absolute error independent of unbounded .
There is no later hidden growth regime. The exact positive system keeps in , because its vector field points inward at every boundary. Its positive equilibrium point is , with Jacobian matrix having trace and determinant . The origin has no attracting direction inside the positive quadrant, while the divergence excludes cycles by the Bendixson-Dulac theorem. The Poincaré-Bendixson theorem therefore gives convergence to the positive stable equilibrium. At fixed small positive , the exact far-field slope is , whose difference from is only relative , consistent with the global leading composite.
For , the polar coordinates satisfy
A positive constant radius must therefore be one. The nonconstant periodic solutions are
with least period and counterclockwise orientation. The origin is also a stationary solution, hence periodic in the degenerate sense of a constant function; it is a point rather than a nontrivial closed orbit. There are no other constant-radius solutions.
For , put . Then , a logistic differential equation. Solving this separable differential equation gives
The denominator is positive for every forward time. When , the radius increases monotonically to one; when , it decreases monotonically to one; and when it stays there. Thus every nonzero orbit approaches the unit circle while continuing to rotate. More precisely, its distance from the phase-matched circular solution tends to zero; its Cartesian coordinates do not converge to a single point. For , the solution stays at the origin.
The origin is the only equilibrium point. Its Jacobian matrix is
whose eigenvalues are . It is an unstable focus. The unit circle is an attracting limit cycle, rather than a curve of equilibria: on it the vector field remains nonzero. Linearizing the radial equation at gives , confirming radial attraction. All other radii are strictly monotone, so there are no additional nontrivial periodic orbits.
Figure 1.
Counterclockwise spirals approach an attracting unit circle around an unstable equilibrium
. Details.
At the unscaled state , write the Lévy kernel of a Markov jump process on jump displacements as
The destination-kernel convention translates each Dirac mass by . The total holding rate is the mass of this kernel. At either zero population the corresponding death rates vanish, so no negative states are reached. Births in the total population are bounded by a Yule process with per-individual rate , while deaths only decrease it; this also gives nonexplosion.
Put . This is a density-dependent Markov jump process, with jumps of size and rates of order . Its drift gives the deterministic approximation
Uniqueness preserves the diagonal . For positive , the resulting logistic differential equation has solution
If , it decreases from one to ; if , it increases to that same equilibrium; equality gives the constant solution one. In particular positive rates do not give extinction on this exactly symmetric deterministic trajectory. Transverse perturbations behave differently: subtracting the equations gives , so the positive symmetric equilibrium has an unstable difference direction. The symmetric competition model distinguishes diagonal logistic relaxation from this symmetry-breaking instability.
For an explicit error estimate, use the martingale decomposition of a density-dependent jump process:
Fix and choose . Stop at the first exit from the box . On that box, has Lipschitz constant in the supremum norm. Each coordinate of has jumps bounded by and predictable quadratic variation at most
Indeed a coordinate's positive and negative jump rates are and , and the squared jump size is . Independent jump clocks give zero cross-bracket.
The Bernstein bound for bounded-jump martingales gives, for each coordinate,
The Gronwall inequality applied up to gives
An exit from the box already entails an error larger than , by the choice of , so the stopped estimate controls the event of any error on the full interval. Take and use the union bound on the two coordinates:
For fixed time horizon and tolerance this is exponentially small in , proving a fluid limit. The transverse instability explains why this finite-horizon estimate should not be extrapolated to arbitrarily long times without changing its constants.
Write . The admissible equilibrium points are , when , and when . The Jacobian matrix is
At the eigenvalues are and ; at they are and . At a positive coexistence equilibrium point,
It is therefore asymptotically stable. It is a node for , a focus for the reverse inequality, and has repeated real eigenvalues at equality. The node–focus change does not alter the attracting equilibrium point's topology.
There are three open physical parameter regions:
  • : extinction. Only is admissible and it is a sink.
  • , : predator extinction. is a saddle, and is a sink.
  • : coexistence. and are saddles on the boundary, while is a sink.
The axes are invariant. On , decays exponentially; on , prey obey the logistic differential equation. Positivity follows because each right-hand side has its population as a factor. Also is bounded by logistic comparison, while satisfies , so all forward trajectories are bounded. The Dulac function has divergence in the interior, excluding periodic orbits by the Bendixson-Dulac criterion.
In the extinction region, , and then is eventually negative. In the prey-only region, the upper bound implies predator decay and convergence of interior prey to . In the coexistence region the Lyapunov function
has . Its positive-quadrant sublevel sets are compact, and the only invariant subset with also has . The LaSalle invariance principle proves convergence of every interior trajectory to . These facts justify the complete qualitative phase portraits, not just their local arrows.
At , and meet in a transcritical bifurcation; positive prey decay algebraically. At with , and exchange stability. On its centre manifold, gives , so the physically positive side is still attracting. There is no interior Hopf bifurcation since the coexistence trace never vanishes.
Figure 1.
Three continuous predator–prey phase portraits and their parameter regions
. Details.
A Markov jump process relative to is an adapted càdlàg process with piecewise-constant paths and finitely many jumps on each bounded time interval, whose conditional future law given depends only on its current state. For a time-homogeneous process on a countable state space, write for its transition rates and . In state , the holding time is exponential of rate and the next state is with probability ; a zero rate makes the state absorbing. The generator is . Nonexplosion is included here so that the process is defined at every finite time.
Here is a finite-jump, bounded-rate form of the Kurtz fluid limit theorem. For each , let be a chain on a countable state space with jumps at rates , with in a fixed finite subset . Assume the nonnegative are uniformly bounded, admissible jumps remain in the state space, and
is globally Lipschitz with constant . Suppose in probability, and let solve , . Then for every finite ,
Bounded total jump rates ensure nonexplosion; the globally Lipschitz drift gives a unique global ODE solution. These explicit hypotheses suffice for the population example, without needing a more general local version of the theorem.
To prove the result, compensate the jumps of this density-dependent Markov jump process. The drift and the coordinate martingales satisfy
Here is predictable quadratic variation, not the sum of realized squared jumps. The compensation identity follows because each possible increment has intensity , and its squared coordinate increment contributes per unit time. Bounded rates and jump sizes make these martingales square-integrable.
We use the Doob L2 maximal inequality in the explicit form for a scalar square-integrable martingale starting at zero. Applying it coordinatewise and setting gives
Subtract the ODE integral equation. Lipschitz continuity and Gronwall's inequality yield
The first term tends to zero in probability by hypothesis, and the second by its maximal second-moment estimate. More quantitatively, for ,
This proves the stated uniform finite-horizon fluid limit.
For the cells, division replaces one cell by two and increases population by one. At population , the total rate is the sum of the identical cell rates, namely
The state is absorbing. Set ; its jump is with rate , where on . Extending this function by zero outside makes it bounded, nonnegative and globally Lipschitz, with Lipschitz constant one. Starting from , or from when integer rounding is necessary, the limiting ODE is the logistic differential equation
Separation of variables, or differentiating , gives its solution. The logistic cell-division process therefore satisfies
Here , so the maximal martingale second moment is at most . With exact initial density , the preceding pathwise bound even gives . With integer rounding, the bound follows from and . Thus the deterministic logistic curve approximates the density with fluctuations of order on fixed time intervals. The convergence statement is on finite horizons; it does not interchange the limits and .
Use the complete system in the PDF: the TeX omits . Both coordinate axes are invariant, and the equilibrium points are , and . The Jacobian matrix is
At its eigenvalues are , and at they are : both points are saddle equilibria. At the coexistence point,
Thus the interior point is a stable spiral. The nontrivial nullclines are and : increases below the first and decreases above it, while increases to the right of the second and decreases to the left. The rotation near coexistence is counterclockwise.
A local linearization by itself does not prove that every interior solution has the same limit. For this logistic predator-prey model, let and use the Lyapunov function
The inequality makes nonnegative, with equality only at coexistence. Its sublevel sets are compact inside the positive quadrant, because diverges both at the axes and at infinity. Direct differentiation cancels the predator-prey cross terms and gives
On one has , and remaining there requires . The largest invariant subset is therefore the coexistence point. The LaSalle invariance principle proves
On , decays as toward the origin. On , the logistic differential equation takes every to one. The origin itself stays fixed. These boundary trajectories are the exceptions to the interior convergence statement.
For , differentiate and use the angular derivative . The polar coordinates satisfy
Hence the rotation is clockwise. The origin is an unstable equilibrium, while is a limit cycle of period . Writing gives the logistic differential equation . For ,
All nonzero solutions tend to the unit-circle attracting limit cycle as , spiralling outward for and inward for . For the solution exists for every real time and approaches the origin as ; for it remains periodic in both time directions. For , the denominator first vanishes at
This finite-time blow-up of an ordinary differential equation occurs backward in time. Such exterior solutions have no behavior: their maximal time interval is . The zero solution exists for all time and has no defined polar angle.
Put . The shared travelling phase again cancels advection in the nonlinear Ginzburg-Landau equation; the remaining amplitude ordinary differential equation is
For , obeys the logistic differential equation . Equivalently, solves , whence . The nonconstant cubic amplitude saturation solution is
It takes the specified at . The denominator of the cubic amplitude saturation solution remains positive for all future time: if its bracket exceeds one, and if it lies between and one. Thus the positive solution is global forward in time. would give the excluded constant solution; would remain at zero rather than relax to the positive wave.