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Take the original sphere on the tube axis, with diagonal positive hydrodynamic resistance matrix $R_0$ and axial force $W>0$. Add a freely moving neutral sphere and extend its rigid velocity through its interior. The resulting field is admissible in the original domain, so its dissipation obeys $D>q^T{R}_{0}q\ge R_{zz}{V}_z^2$ for any finite nonoverlapping placement of the extra sphere. Since only the first sphere does external work, $D=W{V}_z$, giving the displayed bound without assuming zero transverse velocity or rotation. Strictness follows because equality would require the original nontrivial analytic flow to have zero strain throughout the added ball.

Two widely separated spheres rigidly joined by a hydrodynamically negligible connector obey the free-draining approximation. Their translational hydrodynamic resistance matrix is $6\pi \mu (a_1+a_2)I$. The displayed point is the centre about which every pure translation has zero hydrodynamic torque: ${\sum }_i{\zeta }_i({\mathbf{r}}_i-{\mathbf{r}}_c)=0$. A pure rotation about that point gives no resultant force at this order. This is an exact cancellation within the approximation, rather than an exact formula at arbitrary finite separation. It does not assert that every chiral rigid body has an origin eliminating its translation-rotation coupling.

With a consistent force convention, the mobility matrix maps applied generalized forces to rigid-body velocities; it is the inverse of the hydrodynamic resistance matrix. For a sphere with force on the fluid $\mathbf{F}$, its isolated mobility is $1/(6\pi \mu a)$. Coupling between bodies represents hydrodynamic interactions.

Let $p_T=n{k}_{B}T$, $C=⟨\mathbf{R}{\mathbf{R}}^T⟩$, and $M=I+\alpha \sigma$, so the single-bead mobility is $M/\zeta$. Assume an incompressible solvent, constant $n,H,T,\zeta$, negligible bead inertia, and a symmetric positive-definite $M$. The last requirement makes the resistance invertible and Brownian diffusion physically admissible. At a given position and time, $M$ depends on the moment $C$ but not on the individual configuration $\mathbf{R}$.

Use the source's gradient convention $K_{ij}={\partial }_i{u}_j$, and write $G=K^T$ for the component-first gradient. Then $\mathbf{u}({\mathbf{r}}_2)-\mathbf{u}({\mathbf{r}}_1)\simeq G\mathbf{R}$. The spring forces on the two beads are ${\mathbf{F}}_1=H\mathbf{R}$ and ${\mathbf{F}}_2=-H\mathbf{R}$. Their overdamped probability currents areHere the Brownian term written as $-k_{B}T\nabla \psi$ is a force weighted by probability density. Its corresponding conditional osmotic force is $-k_{B}T\nabla \log \psi$; treating the given term as an unweighted deterministic force per bead would not produce the stated Fokker–Planck equation.

With midpoint $\mathbf{x}=({\mathbf{r}}_1+{\mathbf{r}}_2)/2$, the derivatives are ${\nabla }_{r_1}=\frac{1}{2}{\nabla }_x-{\nabla }_R$ and ${\nabla }_{r_2}=\frac{1}{2}{\nabla }_x+{\nabla }_R$. Subtracting the bead currents gives the configuration currentThe midpoint current is $\mathbf{u}\psi$ after neglecting the specified spatial Brownian diffusion. Configuration probability conservation therefore givesSince $G\mathbf{R}$ has components $R_j{\partial }_j{u}_i$, this agrees with the source's $\mathbf{R}\cdot \nabla \mathbf{u}$. The factor two in both spring relaxation and configuration diffusivity comes from relative motion of both beads.

The second moment $C$ is the unnormalized conformation tensor. Now multiply by $R_i{R}_j$ and integrate over configuration space. Assume sufficiently decaying tails so every boundary term at infinite separation vanishes. The affine current gives $GC+C{G}^T$; spring drift gives $-(2H/\zeta )(MC+CM)$; two integrations by parts of diffusion give $(4k_{B}T/\zeta )M$. HenceBecause $\sigma =nHC-p_{T}I$, $M$ is a polynomial in $C$ and commutes with it. ThusThis is the Hookean dumbbell conformation equation with anisotropic mobility, an exact second-moment closure within the stated model; no Gaussian approximation to $\psi$ was needed.

The upper-convected derivative of the identity is $I^{\nabla }=-(G+G^T)$. Applying it to the polymer stress therefore givesConsequently the coefficients and the closed stress equation arewhere ${\sigma }^{\nabla }=D_t\sigma -\sigma K-K^T\sigma$ in the printed convention. Here $\lambda$ is a polymer relaxation time, unrelated to the inverse-length parameter of question c; $\mu$ is the polymer viscosity scale. The coefficient $\alpha$ has inverse-stress units. In the dimensional mobility parameter in the Giesekus model, the dimensionless factor is $b=\alpha p_T$, not the dimensional $\alpha$ printed in this problem.

The upper-convected rate removes the affine stretching and rotation of material connectors. If $\dot{F}=GF$, then a tensor transported purely as $C=F{C}_0{F}^T$ has $C^{\nabla }=0$. Under a time-dependent rigid rotation $Q$, the rate transforms as $Q{C}^{\nabla }{Q}^T$: this is material frame indifference, not an ordinary componentwise time derivative. The resulting equation is the Giesekus model in these dimensional conventions.

The isotropic subtraction is fixed by zero polymer extra stress at thermal equilibrium. With no flow and positive mobility, the stationary distribution isso $\boxed{{\sigma }_{\mathrm{eq}}=0}$. The constitutive equation correspondingly has a zero-stress quiescent state and a Newtonian low-rate limit $\sigma \simeq \mu (K+K^T)$. Without subtracting exactly $p_{T}I$, the spring contribution would retain an equilibrium isotropic stress and the equation would acquire corresponding reference terms. Because mobility itself depends on $\sigma$, that reference cannot simply be shifted while retaining the same model. The separate fluid pressure enforcing incompressibility is not included in this polymer extra stress.

For steady uniaxial extension of the Giesekus model, the component-first velocity gradient is diagonal, $G=\mathrm{diag}(\gamma ,-\gamma /2,-\gamma /2)$. For a steady spatially uniform diagonal stress, the material derivative vanishes and the convected terms are $-2\gamma {\sigma }_{11}$ and $+\gamma {\sigma }_{22}$. The off-diagonal equations vanish identically, and symmetry permits equal transverse components. The remaining equations areSet $W=\lambda \gamma \ge 0$, $b=\alpha p_T$ and $s_i={\sigma }_{ii}/p_T$, with $\mu =\lambda p_T$. For $b>0$, the branches continuously connected to zero stress at zero rate areA sufficient nondegenerate range for physical solutions at every nonnegative rate is $0<b<1$. Indeed $M=(1-b)I+bA$ with $A=HC/(k_{B}T)$, so positive conformation implies positive mobility. The longitudinal quadratic equation is nonpositive at $s_1=0$ and has a nonnegative root of a polynomial. The transverse quadratic equation has values $b-1<0$ at $s_2=-1$ and $W\ge 0$ at $s_2=0$, so its larger root of a polynomial lies in $(-1,0]$. Thus the displayed branches satisfy $s_i>-1$, giving $A_i=1+s_i>0$. The other algebraic roots do not provide a physical branch connected to the equilibrium distribution. For constant positive diagonal $A$, the associated stationary configuration distribution is a Gaussian with covariance $C=(k_{B}T/H)A$.

For positive $\alpha$ on the physical branch, the stretching stress remains finite through $W=1/2$. At that rate,At large extension rate,Thus the stress continues to grow, but the uniaxial extensional viscosity ${\eta }_E=({\sigma }_{11}-{\sigma }_{22})/\gamma$ approaches the finite plateau $2\lambda /\alpha$ rather than developing a finite-rate pole. The transverse stress approaches $-p_T$ as its conformation tensor variance becomes small. At small rate the two stresses are $2\mu \gamma$ and $-\mu \gamma$, so ${\eta }_E\to 3\mu$.

When $\alpha =0$, the physical steady branch instead isThe stretching stress diverges as $W\to 1/2$ from below. Above that threshold its algebraic continuation has $1+s_1=1/(1-2W)<0$ and cannot be a conformation covariance. There is no physical finite steady stretching state. The transient equationgives linear growth at $W=1/2$ and exponential growth at larger rates, the Hookean coil-stretch failure of the upper-convected Maxwell limit.

Parameter qualification. Positivity of $\alpha$ alone, without a restriction on $b=\alpha n{k}_{B}T$, does not ensure real physical steady solutions for the entire tensor. For example $b=2$, $W=1$ makes the transverse equation $2s_2^2+2s_2+1=0$, which has no real root. Its diffusion/resistance interpretation is already inadmissible for some conformations. The physically positive-mobility regime stated above avoids that counterexample; the source's positive-$\alpha$ case must not be taken as a guarantee for arbitrary parameters.

For two Stokes flows in the same fluid domain with the same dynamic viscosity, symmetry of the Newtonian fluid stress tensor and incompressibility give the local reciprocal identityIn index notation, its remaining cross-gradient terms are both $2\mu e:e^{\prime }$. On the remote sphere the integrated reciprocal flux tends to zero: the velocity is $O(R^{-1})$, the stress tensor is $O(R^{-2})$ and the area is $O(R^2)$. On the body, the no-slip boundary condition makes the two velocities the constants $\mathbf{U},{\mathbf{U}}^{\prime }$. The Lorentz reciprocal theorem therefore givesBy the Linearity of Stokes flow, the sum of the two velocity-pressure fields satisfies the boundary conditions for body velocity $\mathbf{U}+{\mathbf{U}}^{\prime }$. Uniqueness of the exterior Stokes flow identifies it with the third solution. Since the stress tensor and its surface integral are linear too, $\mathbf{F}(\mathbf{U}+{\mathbf{U}}^{\prime })=\mathbf{F}(\mathbf{U})+\mathbf{F}({\mathbf{U}}^{\prime })$; scalar multiplication follows in the same way. Thus $\mathbf{F}=-A\mathbf{U}$ for a fixed-orientation hydrodynamic resistance matrix $A$. Substituting arbitrary $\mathbf{U},{\mathbf{U}}^{\prime }$ in the reciprocal identity yieldsThe minus sign is natural because viscous dissipation gives $-\mathbf{F}\cdot \mathbf{U}=2\mu \int e:edV>0$ for nonzero rigid translation in an otherwise stationary fluid; $A$ is a positive-definite matrix.

Use the free-draining approximation: neglect the hydrodynamic interaction between the widely separated spheres and neglect the connector's viscous drag. Stokes law gives ${\mathbf{f}}_i=-6\pi \mu a_i\mathbf{U}$ for each sphere. Their resultant isThis is the leading two-sphere approximation, not an exact finite-separation hydrodynamic resistance matrix. Writing $d=|{\mathbf{r}}_2-{\mathbf{r}}_1|$ and $\mathbf{n}=({\mathbf{r}}_2-{\mathbf{r}}_1)/d$, the first Stokeslet reflection gives, if the connector remains negligible,The negative correction expresses reduced drag from fluid entrainment by the other sphere. A very thin but arbitrarily long rod need not have negligible total drag: its typical transverse drag scales as $\mu d/\log (d/b)$ for rod radius $b$. Thus the simple boxed result uses the intended negligibly dragging connector limit as well as $a_i/d\ll 1$; the two printed size inequalities alone do not specify a uniform limit in which rod drag vanishes.

Put $r=|\mathbf{x}|$ and $s=\mathbf{F}\cdot \mathbf{x}$. In the Unscaled Papkovich–Neuber representation, take the harmonic potentials $\mathbf{\Phi }=-\mathbf{F}/(8\pi \mu r)$ and $\chi =0$. Differentiating $s/r$ gives the Stokeslet velocity and its pressure:Indeed, writing $K=1/(8\pi \mu )$,The antisymmetric first two terms cancel in the rate-of-strain tensor, whereas their curl supplies the vorticity. ThusThe rate-of-strain tensor has zero trace, as required by incompressibility. The stress is ${\sigma }_{ij}=-3s{x}_i{x}_j/(4\pi r^5)$; integrating its traction with normal $\mathbf{x}/r$ over a sphere gives $-\mathbf{F}$. This fixes the force normalization of the Stokeslet.

For a sphere in a uniform straining Stokes flow, write the ambient velocity as $\mathbf{Ex}$, where $\mathbf{E}$ is symmetric and trace-free, and set $q=\mathbf{x}\cdot \mathbf{Ex}$. Symmetry gives zero translation and rotation of the centred force-free, torque-free sphere. Seek a decaying disturbance using the Unscaled Papkovich–Neuber representation withThe components of $\mathbf{Ex}/r^3$ are derivatives of $1/r$, and $q/r^5$ is a trace-free harmonic quadrupole, so both potentials satisfy Laplace's equation outside the sphere. Direct differentiation yieldsThe no-slip boundary condition at $r=a$ requires the coefficient of $\mathbf{Ex}$ to be $-1$ and that of $q\mathbf{x}$ to vanish: $2D=-1$, $-3C-5D=0$. Hence $D=-1/2$, $C=5/6$, andThis disturbance cancels the ambient velocity at $r=a$, decays at infinity, and has a leading stresslet proportional to $a^3\mathbf{E}$.

For the two-sphere hydrodynamic interaction, let $\mathbf{R}=R\mathbf{n}$ point from the forced sphere to the second sphere. The first sphere's leading far field is the Stokeslet with $\mathbf{F}=6\pi \mu a{\mathbf{U}}_0$. Its incident rate-of-strain tensor at the second sphere isThe force-free sphere translates with the local uniform incident flow, and the torque-free sphere rotates with half the incident vorticity. Those parts cause no leading Stokeslet or rotlet disturbance. The remaining incident strain produces the stresslet just calculated. At displacement $-\mathbf{R}$ from the second sphere its velocity isThe first sphere translates in this returned incident flow. Finite-size sampling and higher incident multipoles enter only beyond the retained reflection order. Thus the stresslet reflection between two force-free and forced spheres givesThe factor $|{\mathbf{U}}_0|$ makes explicit the velocity scale implicit in the printed remainder.

The resulting input power, equal to the viscous dissipation, isIt decreases at fixed force. The Minimum-dissipation theorem for Stokes flow instead compares admissible fields at fixed first-sphere velocity: extending a two-sphere field rigidly into the second sphere gives an admissible one-sphere field, so the extra rigid inclusion can only increase the minimum resistance. Allowing the second sphere to choose its translation and rotation minimizes over its rigid motions, producing zero force and couple there. An increase of this effective hydrodynamic resistance matrix decreases its inverse hydrodynamic mobility matrix, and hence decreases fixed-force power. The negative mobility correction is therefore consistent with minimum dissipation.

If the second sphere is prevented from rotating, its angular velocity relative to the incident fluid becomesUse the rotating sphere in Stokes flow at $-\mathbf{R}$. The additional reflected rotlet givesAdding this rotation-clamped reflection between two spheres to the strain reflection,The holding couple supplies no power because the held sphere's angular velocity is zero.

Let downward velocity be positive in this part, and let $W>0$ be the dense sphere's weight minus buoyancy. For the original centreline configuration, write its six-component translation-and-rotation velocity as $q$, and its diagonal positive hydrodynamic resistance matrix as $R_0$. Its single-sphere dissipation isWith only axial force and no torque, the original velocity is $V_0=W/R_{zz}$.

Now add the neutral sphere. In its actual Stokes flow, extend the velocity rigidly through the new sphere's interior. This extension is a valid incompressible trial field for the domain without that sphere, with exactly the dense sphere's actual six boundary velocities; its strain and dissipation inside the inserted ball are zero. The Minimum-dissipation theorem for Stokes flow and extra dissipation due to a rigid inclusion therefore giveThe neutral sphere has zero net external force and torque, and the dense sphere has only the axial force $W$. The dissipation equals total external power, so $D=W{V}_z$. Positivity gives $V_z>0$, and the comparison yields $V_z\le W/R_{zz}$ without assuming that either sphere translates only vertically or that either angular velocity vanishes.

For a finite nonoverlapping placement, the first inequality is strict. Equality would make the extended field the old Stokes solution and thus make that solution rigid throughout the added ball. Analyticity and connectedness then force its strain to vanish throughout the original fluid domain; the stationary tube wall and quiescent far field would force the whole motion to vanish, contradicting $V_z>0$. Hence the force-free inclusion reduces axial mobility of a centred settling sphere:The difference tends to zero as the added sphere recedes infinitely far away. The lateral and rotational contributions are nonnegative in the resistance comparison, which is precisely why they cannot invalidate the bound.

For two Stokes flows $(u^{(1)},{\sigma }^{(1)})$ and $(u^{(2)},{\sigma }^{(2)})$ of the same dynamic viscosity in the same region, with no volume force, the Lorentz reciprocal theorem statesIndeed, the divergence of their cross-work difference isThe pressure terms vanish by incompressible flow, and the derivatives of the Cauchy stress tensors vanish by the Stokes equation. The divergence theorem proves the result.

Use the convention that $F,G$ are the force and torque exerted by the body on the fluid. By Linearity of Stokes flow, $(F,G)=\mathsf{R}(U,\Omega )$. Applying the Lorentz reciprocal theorem to two independent rigid motions gives $q_1^T\mathsf{R}{q}_2=q_2^T\mathsf{R}{q}_1$, so the hydrodynamic resistance matrix is a symmetric matrix. The boundary-power identity givesThis is strictly positive for nonzero rigid motion: equality would imply $e=0$, hence a rigid motion throughout the connected fluid, which must vanish since the fluid is at rest at infinity. The no-slip boundary condition would then force $q=0$. Therefore $\mathsf{R}$ is a positive-definite matrix. Reversing to the fluid-on-body force changes the sign of the force law, not the positive resistance coefficients.

For the two rods, take the torque about $O$ and use body axes. On the $x$-rod, $X=(s,0,0)$, $0\le s\le 2L$, and its slender-body force density isOn the $y$-rod it isIntegrate $f$ and $X\times f$ along both rods, using $\int ds=2L$, $\int sds=2L^2$ and $\int s^{2}ds=8L^3/3$. A convenient dimensionally uniform statement of the complete right-angle two-rod resistance matrix isIn physical coordinates this means translation–translation entries scale as $CL$, the two translation–rotation blocks as $C{L}^2$, and rotation–rotation entries as $C{L}^3$. The planar block has inverseThis verifies the printed hydrodynamic mobility matrix with its third velocity component $L{\Omega }_z$; the TeX aid's ${\Omega }_x$ is an OCR error.

Take laboratory vertical velocity positive upwards. The body axes are $e_1=(\cos \theta ,\sin \theta )$ and $e_2=(-\sin \theta ,\cos \theta )$. In quasistatic sedimentation the force and torque exerted on the fluid equal the gravitational resultants on the body:The out-of-plane block is unforced, so its positive hydrodynamic resistance matrix gives $U_z={\Omega }_x={\Omega }_y=0$. Substitution in the planar hydrodynamic mobility matrix givesandFor $0\le \lambda <1$, this ordinary differential equation has positive right side on $0\le \theta <\pi /4$, with a stable zero at $\pi /4$. Uniqueness prevents crossing that equilibrium in finite time. For $\lambda >1$ the initial angular velocity is negative, and the stable equilibrium reached from zero is $-3\pi /4$. The heavier-end body turns clockwise through $3\pi /4$; it does not settle at $\pi /4$. More explicitly, with $K=(1-\lambda )mg/(4C{L}^2)$,where the continuous branch has $\theta -\pi /4\in (-\pi ,0)$. At $\lambda =1$, $\theta$ remains zero.

Transforming the translational velocity back to laboratory axes givesFor $\lambda \ne 1$, eliminate time between $U_h$ and the angular velocity:At either limiting orientation $\sin \theta =\cos \theta$. Thus both cases have the same net horizontal displacement,For $\lambda <1$ the drift is monotonically rightwards. For $\lambda >1$ it first moves left, reaching $x_O-x_O(0)=(2L/3)(1-\sqrt{2})$ at $\theta =-\pi /4$, then reverses. The total horizontal path length in this second case is $(2L/3)(2\sqrt{2}-1)$, whereas its net displacement is $2L/3$ rightwards. At $\lambda =1$ it falls vertically without rotation or drift; taking the infinite-time limit before $\lambda \to 1$ is consequently singular.

For the requested sedimentation drift of a weighted two-rod body sketch, a full parametric trajectory follows by also integrating $U_v/\dot{\theta }$. Put $\psi =\theta -\pi /4$ and set the initial height to zero:It tends to $-\infty$ in either case while $x_O$ tends to $2L/3$. At $\lambda <1$ both rods end pointing upwards from $O$ symmetrically; at $\lambda >1$ they end pointing downwards symmetrically. The asymptotic downward speed is $(1+2\lambda )mg/(3CL)$.

Use the same resistive-force theory convention as before: the force density is the fluid's force on the filament. Let $\phi$ be the usual cylindrical azimuth. Choose the orientation of a left-handed helix so that, as $z$ increases, $\phi$ decreases. With $s=\sin \theta$, $c=\cos \theta$,Reversing the orientation of the tangent leaves the local drag tensor unchanged. Define $k={\xi }_{\perp }-{\xi }_{\parallel }$. From resistive-force theory, the relevant force-density components areThese scalar axial and azimuthal components are uniform along the helix. Integration over its contour length $\ell$, and use of the axial torque density $a{f}_{\phi }$, give the axial resistance matrix of a slender helix:Thus $B=C$, as also required by the Lorentz reciprocal theorem. The signs here belong to $(F,L{)}^T=-\left(\begin{array}{cc}A& B\\ B& D\end{array}\right)(U,\Omega {)}^T$, with $F,L$ forces and moments on the helix. The handedness reversal of helical hydrodynamic resistance reverses $B,C$ but leaves $A,D$ unchanged. If the positive rotation direction or handedness convention is reversed, the coupling sign reverses with it.

As a check on physical admissibility, the determinant of helical resistance in resistive-force theory isTogether with $A,D>0$, this makes the hydrodynamic resistance matrix positive definite and the viscous power loss $A{U}^2+2BU\Omega +D{\Omega }^2$ positive. Isotropic local drag would have $B=0$, so rotating a helix would not propel it in this approximation.

For the pair, use a local additive resistive-force theory model: both helices have the same drag coefficients, and inter-helix hydrodynamic interactions and unresolved end effects are omitted. The right-handed helix has resistance entries $(nA,-nB,-nB,nD)$. Define the signed motor motion byThe printed relative rotation specifies a magnitude rather than which helix rotates relative to which. Choosing ${\Omega }_2-{\Omega }_1=\omega$ instead reverses all signed speeds below.

The common translation speed $U$ and zero total force and torque obeyInternal motor forces and torques cancel in these totals. IntroduceSolving the three linear equations yields the opposite-handed counterrotating helical swimmer:For $n>0$, ${\xi }_{\perp }>{\xi }_{\parallel }$ and $\omega >0$, $B>0$, so $U<0$, ${\Omega }_1>0$ and ${\Omega }_2<0$. The opposite-handed helices counterrotate but contribute thrust in the same axial direction. The formula satisfies the specified relative rotation without identifying either laboratory rotation rate with the motor rate.

At $n=0$, the second helix supplies no hydrodynamic resistance. The remaining helix must have $F=L=0$, and invertibility of its hydrodynamic resistance matrix forces $U={\Omega }_1=0$. Formally ${\Omega }_2=-\omega$ is the rotation of a zero-resistance motor shaft or vanishing second rotor; there is no physical finite second helix to propel or to provide a reaction torque. This is the vanishing reaction rotor in a helical swimmer.

At $n=1$, equal-length opposite-handed helices haveThe translation-generated axial torques cancel between the two helices, and equal counterrotation balances the rotational torques. Their propulsive forces add, rather than cancel, because handedness and rotation both reverse. These are equal-length opposite-handed helices.

For $n\to \infty$, the large reaction helix limit isThe increasingly long second helix nearly anchors the whole assembly: its very small translation and rotation suffice to balance the finite force and torque generated by the first helix. The first helix rotates at almost the full motor rate, but moving the large resistive second helix makes the common translation tend to zero. These are idealized limits of the additive local model, rather than a resolution of short-helix end effects or infinitely extended interacting filaments.

Take $\mathbf{F},\mathbf{G}$ to be the force and couple exerted by the body on the fluid, equivalently the external force and couple needed to maintain its motion. This fixes the sign for a positive hydrodynamic resistance matrix. If one uses the fluid's force on the body, both resultants have the opposite sign.

For two Stokes flows $(\mathbf{u},\mathbf{\sigma })$ and $(\tilde{\mathbf{u}},\tilde{\mathbf{\sigma }})$ in the same fluid domain $\mathcal{D}$, with zero body force, the Lorentz reciprocal theorem for Stokes flow isHere $\mathbf{n}$ points out of the fluid. To see the identity, take the divergence of the difference of the two cross-work fluxes. The stress divergences vanish, while incompressibility and symmetry of the Newtonian fluid stress tensor reduce the remaining terms to $2\mu (\mathbf{e}:\tilde{\mathbf{e}}-\tilde{\mathbf{e}}:\mathbf{e})=0$. The divergence theorem proves the result.

On the body, the no-slip boundary condition is $\mathbf{u}=\mathbf{U}+\mathbf{\Omega }\times \mathbf{r}$, and the far-field contribution vanishes for decaying exterior flows. Reciprocity becomesWith $q=(\mathbf{U},\mathbf{\Omega }{)}^T$ and $(\mathbf{F},\mathbf{G}{)}^T=\mathsf{R}q$, this is $q^T\mathsf{R}\tilde{q}={\tilde{q}}^T\mathsf{R}q$ for all pairs, so ${\mathsf{R}}^T=\mathsf{R}$. The power identity givesEquality would force zero strain throughout the connected exterior domain, hence a rigid fluid motion. Decay at infinity makes that motion zero, and the no-slip boundary condition then gives $q=0$. This proves the symmetry and positivity of a rigid-body resistance matrix:

For the helix, assume $0<\varphi <\pi /2$ and use cylindrical unit vectors. Its arclength element and tangent areIntegrating to $\theta =L\cos \varphi /b$ gives $\boxed{\text{total arc length}=L}$. In a combined axial translation and rotation, $\mathbf{V}=U{\mathbf{e}}_z+b\Omega {\mathbf{e}}_{\theta }$. The slender-body force density, or local resistive-force theory, givesWriting $s_{\varphi }=\sin \varphi$, $c_{\varphi }=\cos \varphi$, its axial and azimuthal components areThe axial couple density is $b{f}_{\theta }$. Integration over arclength gives the axial resistance matrix of a slender helix:Thus pure translation gives $(F_z,G_z)=(AU,BU)$, and pure rotation gives $(F_z,G_z)=(B\Omega ,D\Omega )$. The mixed coefficients agree, as reciprocity requires, and $AD-B^2=C^2{L}^2{b}^2/2>0$. The sign of $B$ follows the handedness in the parametrization; reversing handedness reverses propulsion.

For the helical microswimmer with a spherical head, the diagram's rotation relation is ${\Omega }_0=\Omega -\omega$. Neutral buoyancy and the absence of external forcing make the whole swimmer force-free and torque-free. Neglecting interactions between its parts givesWith $\Delta =(A_0+A)(D_0+D)-B^2>0$, solving this pair givesThe head counterrotates, providing the reaction to the flagellum's rotation. The figure fixes this relative rotation convention; it is duplicated and corrupted in the local TeX.

For a very large head, the coefficient regime is $A_0\gg A$ and $D_0\gg D$. ConsequentlyThe large translational resistance $A_0$ makes the microswimmer slow even though the flagellum rotates almost at the motor rate.

For a very small head, $A_0\ll A$ and $D_0\ll D$. The appropriate denominator retains the translation-rotation coupling:Both tend to zero with $D_0\propto a^3$, while ${\Omega }_0\simeq -\omega$. The motor mainly rotates the low-resistance head, and produces little flagellar motion relative to the fluid.

In the intermediate coefficient regime $A_0\ll A$ and $D_0\gg D$, the head supplies a strong rotational reaction with little translational penalty. ThenThe head radius has dropped out. To find the optimal pitch of a helical microswimmer, put $z=\tan \varphi$. The dimensionless speed becomes $z/(z^2+2)$ and has derivative $(2-z^2)/(z^2+2{)}^2$. Hence, for $\omega >0$ with the chosen handedness,The angle is measured from the horizontal plane. An intermediate pitch combines axial and transverse tangent directions, allowing anisotropic drag to convert rotation into translation; either extreme removes that coupling.

The geometric head-size regimes quoted in the paper hold with the logarithmic slenderness factor treated as fixed. More precisely, $A_0\ll A$ requires $a\ll L(1+{\cos }^2\varphi )/(3|\log ϵ|)$, while $D_0\gg D$ requires $a^3\gg L{b}^2(1+{\sin }^2\varphi )/(4|\log ϵ|)$. These coefficient inequalities state the validity of the intermediate approximation when the logarithm is quantitatively important. The many-turn and local-drag assumptions of slender-body theory also remain in force.