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A capture-recapture model is a statistical model for repeated observations of identifiable individuals, including those missed on some occasions. A closed-population capture-recapture model estimates a fixed population cardinality; models allowing individuals to enter or leave can instead describe survival and recruitment. The sampling assumptions determine which likelihood function is appropriate.

For the two-sample closed-population capture-recapture model, condition on the sample sizes $M$ and $C$. If the second sample selects $C$ distinct individuals from $N$ by simple random sampling, then the overlap $R$ has the hypergeometric distribution:Here $r$ is an integer and the probability mass function is zero outside these bounds. The binomial coefficients count choices of $r$ marked and $C-r$ unmarked individuals, divided by all equally likely size-$C$ subsets. The hypergeometric distribution has expected valueEquating the observed overlap with this expected value gives the Lincoln-Petersen estimator. The underlying sampling without replacement, probability mass function, and expected value are developed in Raymond P. Guiteras, MIT 14.30 hypergeometric distribution handout, §§1–4 and 8.

For the toy counts $M=20$, $C=25$, and $R=5$, hold the observations fixed and vary the unknown integer $N$. The likelihood function isValues below forty cannot contain the observed set union. Exact comparison of consecutive likelihoods gives two tied maximum-likelihood estimates, $N=99$ and $N=100$. The Lincoln-Petersen estimator returns $100$; the Chapman estimator returns $90$ and is a different rule for estimation.

The vertical scale is $L(N)/{\max }_{n\ge 40}L(n)$. This relative likelihood compares how well candidate statistical parameters explain the same observations; it is not a posterior probability. The displayed range $40\le N\le 300$ is only a plotting window. The likelihood function remains positive for every integer $N\ge 40$ and tends to zero as $N$ increases, so the curve has a continuing right tail. The data do not justify truncating statistical inference at three hundred. The conditional hypergeometric distribution formulation also appears in Louis-Paul Rivest and Mamadou Yauck, Small Sample Estimators for Two-way Capture Recapture Experiments, §3, equation (5).

Zero recaptures require care. With $R=0$ and positive $M,C$, the Lincoln-Petersen estimator has no finite ratio, but the capture-recapture model still gives a valid likelihood function:This likelihood function increases toward one as $N$ grows, so there is no finite maximum-likelihood estimate. For the same sample sizes twenty and twenty-five with zero overlap, forty-five distinct individuals would be observed. Lack of overlap supplies a lower bound but very weak information about large population cardinalities.

In more elaborate capture-recapture models, a first-recapture array $m_{ij}$ counts individuals released at occasion $i$ and next observed at $j$. A probability of survival ${\varphi }_j$ and a probability of detection $P_{j+1}$ contribute a no-observation continuation factor ${\varphi }_j(1-P_{j+1})$. The resulting likelihood function depends on both survival and detection; these probabilities need not be separately identifiable at the last observation interval. Missing-count EM for capture-recapture describes one way to fit such a model, and identifiability remains a separate issue from computing an estimator output.

For a real random variable $X$ with finite variance ${\sigma }^2$, expected value $\mu$, and a threshold $a>0$, Chebyshev's inequality givesOne can also cap this upper bound at one, since a probability cannot exceed one. If $\sigma >0$, taking $a=k\sigma$ gives $\mathbb{P}(|X-\mu |\ge k\sigma )\le 1/k^2$. For example, at least $3/4$ of the probability lies strictly within two standard deviations of the mean. This statement needs only the finite variance assumption. If the variance is zero, $X=\mu$ almost surely and every positive deviation threshold has zero tail probability.

Consider a random variable uniformly distributed on $\{-2,-1,0,1,2\}$. Its expected value is zero and its variance is $(4+1+0+1+4)/5=2$. At $a=2$ the actual tail probability is $2/5$, while Chebyshev's inequality gives $2/2^2=1/2$. At $a=3$ the actual tail probability is zero, while the bound is $2/9$. A valid bound need not describe the exact shape or bounded support of a particular probability distribution.

For the standard normal distribution, the actual tail probabilities beyond two and three standard deviations are approximately $0.0455003$ and $0.0026998$, whereas the general bounds are $1/4$ and $1/9$. More knowledge of the probability distribution can therefore improve the estimate. The comparison does not diminish the guarantee: the three-point example in equality in Chebyshev's inequality attains it exactly.

The proof is a one-line use of Markov's inequality on the nonnegative random variable $Y=(X-\mu {)}^2$. The event $|X-\mu |\ge a$ equals $Y\ge a^2$, and $\mathbb{E}Y={\sigma }^2$. This explains why the denominator is squared and why a finite variance supplies the necessary information. ^[ref].

One important use is control of a sample mean. For independent and identically distributed random variables with finite variance ${\sigma }^2$, the variance of a sample mean is ${\sigma }^2/n$. HenceFor every fixed tolerance, the bound tends to zero as the sample size grows, proving the finite-variance case of the weak law of large numbers. For ${\sigma }^2=1$ and $\epsilon =0.1$, $n=2000$ makes the bound $0.05$. This is a sufficient bound under the stated assumptions, rather than an optimal required sample size.

An early historical source is Pafnuty Chebyshev's 1867 paper “Des valeurs moyennes”, published in French translation by N. de Khanikof. Its opening theorem bounds deviations of sums using their expected values and second moments, linking averages with control of exceptional outcomes. ^[ref].

The coverage probability is the repeated-sampling probability that a random confidence interval contains the fixed true statistical parameter. A finite-sample level-$1-\alpha$ procedure has coverage probability at least $1-\alpha$ at every admissible statistical parameter; exact nominal coverage means equality. An asymptotically calibrated procedure approaches the nominal value as sample size grows, under its stated assumptions. Higher-order Edgeworth expansions can quantify the discrepancy from nominal coverage probability when their regularity conditions hold.

Observed coverage in finitely many repetitions is itself random. For $m$ independent repetitions of the exact normal mean confidence interval with known variance, the count $K$ of covering confidence intervals has binomial distribution $\mathrm{Binomial}(m,0.95)$. At $m=40$, its expectation is $38$, butConsequently, the 39 successes in the figure are compatible with exact 95% coverage probability. The guarantee follows from the pivotal quantity's probability distribution; the finite observed fraction is an illustration. Also distinguish a confidence interval for a fixed mean from a prediction interval for a future observation: the latter must accommodate the new observation's variation as well as estimation uncertainty.

The design effect is a sampling variance under a complex design divided by the corresponding simple-random-sampling variance. For independent equal-size clusters of size $m$ with exchangeable intraclass correlation $\rho$, a common mean-estimation approximation is $1+(m-1)\rho$. It expresses the loss of effective sample size from positive correlation; finite-population and unequal-cluster-size designs require their own calculation.

A sample size calculation for a binary endpoint requires its own baseline probability, not merely the rate of a related outcome. A fatal-event probability equals the probability of the underlying event times its conditional fatality probability; knowing only that product does not identify the incidence of all events. Thus a mortality rate can size a mortality trial but cannot determine the size for all fatal and nonfatal events without further assumptions. The significance level, allocation, target statistical power and effect definition must also be specified.

For two independent binomial counts, form the Fisher exact test rejection region from their conditional null hypergeometric distribution. Summing alternative joint binomial probabilities over that region gives its exact power. For equal group sizes, the null allocation is symmetric, so the two-sided probability-ordering test agrees with doubled lower-tail probabilities. Conservative test discreteness can make a normal-approximation sample size underpowered even when both samples are large but event counts are low.

For uniform sampling without replacement from a population of $n>1$ objects, the finite population correction accounts for the reduced spread caused by drawing distinct objects. For the marked-object count $K$ in a hypergeometric distribution, put $p=c/n$. ThenRelative to the corresponding binomial distribution, the variance is multiplied by $f^2=(n-r)/(n-1)$ and the standard deviation by $f$. The square-root convention for the correction is used in Philip Stark's discussion of sample sums.^[ref]

The factor has a direct interpretation. For $r=1$, replacement makes no difference and $f=1$. For $r=n$, the entire population is observed and the marked count is exactly $c$, so $f=0$. In the ten-object, five-draw example, $f^2=5/9$ and $(6/5)(5/9)=2/3$. The exceptional population size $n=1$ is deterministic and should be handled directly rather than inserting it into a zero denominator.

A short explanation uses an indicator random variable $I_j$ for a marked object at draw $j$. Each has expected value $p$ and variance $p(1-p)$. For distinct draw positions $i,j$,Since $K={\sum }_{j=1}^r{I}_j$, the variance of a sum includes $r$ individual terms and $r(r-1)$ ordered covariance terms, giving the displayed correction. Linearity of expectation still gives $rp$ despite this dependence.

For arbitrary fixed real population values, use $N>1$ for population size and $1\le n\le N$ for sample size. Write the finite population mean $\bar{Y}=N^{-1}{\sum }_i{y}_i$, the finite population variance $S^2=(N-1{)}^{-1}{\sum }_i(y_i-\bar{Y}{)}^2$ and ${\sigma }_N^2=N^{-1}{\sum }_i(y_i-\bar{Y}{)}^2$. Under simple random sampling without replacement, the sample mean satisfiesThus $1-n/N$ is the variance factor relative to $S^2/n$, while $(N-n)/(N-1)$ is the exact variance ratio to independent uniform sampling with replacement, whose sample mean has variance ${\sigma }_N^2/n$. The standard error uses the square root of the corresponding factor. These formulas agree because ${\sigma }_N^2=(N-1)S^2/N$; the denominator convention must be specified. The sample mean under simple random sampling gives a derivation and an exact six-value example.

A specified test for germination, recording the endpoint, conditions and observation window. For a proposed red/far-red light test, use randomization to allocate one seed lot to darkness, red, red then far-red, and red then far-red then red. Match hydration, temperature, handling and elapsed time before counting; use sham waiting periods to equalize schedules. Use dish replication, retain the dark scientific control, and record radicle emergence using an observation procedure that does not itself provide an uncontrolled light treatment. This is a proposed design, not a claim about the 1952 paper's randomization or replication.

As an original hypothetical counting example, begin with $N=40$ seeds in one dish. Suppose 12 first meet the radicle endpoint by 24 hours, eight additional seeds by 48 hours, and four additional seeds by 72 hours. These invented event counts giveand cumulative sample proportions $\hat{G}(24)=0.30$, $\hat{G}(48)=0.50$ and $\hat{G}(72)=0.60$. The 48-hour increment is $8/40=0.20$, not $0.50$; the latter includes earlier events. Counting the same emerged radicle again at each inspection would inflate the total. These numbers illustrate bookkeeping, not biological rates or treatment effects.

With a fixed denominator and irreversible classification of the first endpoint event, the set counted at an earlier inspection is a subset of the set counted later. Hence $0\le \hat{G}(t)\le 1$ and $\hat{G}(t)$ cannot decrease. This short proof applies to cumulative germination counts even if a later seedling dies. A count of currently surviving seedlings has a different definition and can decrease.

For treatment comparisons, the dish may be the experimental unit: seeds in the same dish share conditions. Forty seeds in one dish do not supply forty independent dish replicates. Report the sample size, replication, endpoint time and denominator; a sample proportion by a deadline measures this assay's outcome, not an unconditional fraction with seed viability.

For a fixed positive sample size $n$ of independent and identically distributed Bernoulli random variables, letThe Hoeffding inequality makes $C_n$ a conservative confidence interval for $p$: ${\mathbb{P}}_p(p\in C_n)\ge 1-\delta$ for every $p\in [0,1]$. Indeed, exclusion from the closed interval implies $|{\hat{p}}_n-p|>r_n$, an event contained in the deviation event bounded by $\delta$. Clipping to $[0,1]$ retains coverage because $p$ already lies there. For $n=738$, $\delta =0.05$, and an observed frequency $369/738=1/2$, $r_n\approx 0.0499924$, giving approximately $[0.450008,0.549992]$.

Coverage concerns repeated independent samples: the unknown $p$ is fixed and the confidence interval is random. This fixed-$n$ guarantee does not justify repeatedly checking the same bound and stopping as soon as it looks favorable. The Hoeffding sample size for Bernoulli frequencies chooses $n$ in advance; guarantees across repeated inspection times require separate control of the additional events, for example through a union bound with an allocated failure probability at each time.

Let $B_1,\dots ,B_n$ be independent and identically distributed Bernoulli random variables with success probability $p$, and let ${\hat{p}}_n=n^{-1}{\sum }_i{B}_i$ be their observed frequency, a sample mean. Since every interval width is one, $V=n$. Substituting $t=n\epsilon$ into the Hoeffding inequality givesFor tolerance $0<\epsilon <1$ and failure probability $0<\delta <1$, solving $2e^{-2n{\epsilon }^2}\le \delta$ yields the sufficient sample sizeHere $\log$ is the natural logarithm. For $\epsilon =0.05$ and $\delta =0.05$, $n=738$ guarantees an error smaller than $0.05$ with probability at least $0.95$, for every $p\in [0,1]$. The two-sided bound at this sample size is approximately $0.0499440$. A specified single direction instead needs $n\ge \lceil \log (1/\delta )/(2{\epsilon }^2)\rceil$, or $600$ for these choices. Halving the tolerance multiplies the unrounded sufficient sample size by four; reducing the failure probability tenfold adds $\log (10)/(2{\epsilon }^2)$ before rounding.

This is a conservative sufficient sample size, not a necessary minimum. For a specified $p$, the count $K={\sum }_i{B}_i$ has a binomial distribution, whose exact upper tail probability isFor $k>np$, the Hoeffding inequality bounds this by $\exp [-2(k-np{)}^2/n]$. An arbitrary frequency threshold uses $k=\lceil n(p+\epsilon )\rceil$; if $k>n$, the event is impossible. The discreteness of the count and its variance $np(1-p)$ can make the exact tail probability substantially smaller. At $n=100$, $p=1/2$, $k=60$, the exact upper tail probability is about $0.0284440$, while the bound is $e^{-2}\approx 0.135335$. At $p=1/10$, $k=20$, the same deviation $\epsilon =0.1$ gives the same bound but an exact upper tail probability about $0.00197856$.

An inclusion probability is the probability that a particular unit belongs to the selected sample. For sample size $n$ in simple random sampling from $N$ labels it is $n/N$. It differs from the selection probability at a particular draw position.

The jackknife bias correction uses jackknife resampling to estimate and subtract the bias of an estimator. Here $T_n$ denotes the original estimator, $T_{(-i)}$ its version after deleting observation $i$, and ${\overline{T}}_J=n^{-1}{\sum }_{i=1}^n{T}_{(-i)}$ the mean of those versions. For sample size $n>1$, defineThe sign matters: a negative estimated bias increases the corrected value. The original estimator and all deletion versions must target the same estimand and use the same estimation rule at their respective sample sizes. The definition alone does not make ${\tilde{T}}_J$ an unbiased estimator.

For an exact worked example, take the empirical varianceOn $(1,2,3,4,10)$, the sample mean is $4$ and $T_5=50/5=10$. Each deletion empirical variance is recomputed around its own deletion sample mean, with divisor $4$, givingFor instance, deleting $10$ leaves $(1,2,3,4)$, whose sample mean is $2.5$ and whose empirical variance is $1.25$. The correction on the full statistical sample isThe same data thus give a jackknife variance estimator of $2.5$ for the sample mean, and a corrected empirical variance of $12.5$; these are different estimands.

The exact equality with the usual sample variance is not an accident of these numbers. For any data with $n>1$, write $Q={\sum }_i(x_i-\overline{x}{)}^2=n{T}_n$. Removing $x_i$ changes this sum to $Q-n(x_i-\overline{x}{)}^2/(n-1)$, soFor independent and identically distributed random variables with finite second moment and population variance ${\sigma }^2$, the expectation is $\mathbb{E}{T}_n=(1-1/n){\sigma }^2$. Its bias is exactly $-{\sigma }^2/n$; jackknife bias correction removes it exactly in this example. Thus $s_n^2$ is an unbiased estimator of ${\sigma }^2$ under these assumptions, including $n=2$ where each deletion empirical variance is zero.

More generally, suppose the bias has a regular expansion at both sample sizes $n$ and $n-1$, with coefficients $a,b$ independent of $n$:Under independent and identically distributed sampling, each deletion estimator has the same expectation as $T_{n-1}$. Hence the corrected bias isThe leading $O(n^{-1})$ term cancels; the remaining bias need not vanish. The $O$ notation is big O notation. This argument presumes the expansion, rather than proving it for every statistic; the regularity conditions discussed for the jackknife variance estimator and statistical dependence still require care. Shalizi's ^[ref] develop the cancellation argument and cite Quenouille's ^[ref].

The delete-one form of jackknife resampling recalculates a statistic after deleting each observation in turn. It uses one observed statistical sample to investigate how sensitive an estimator is to its individual observations, and supplies both a jackknife variance estimator and jackknife bias correction. The calculations are deterministic once the data are fixed: no random draws are needed.

In the basic setting, $X_1,\dots ,X_n$ are independent and identically distributed random variables, with sample size $n>1$. Let $x_1,\dots ,x_n$ be the observed values and ${\hat{\theta }}_n=t_n(x_1,\dots ,x_n)$ a scalar estimator of an estimand $\theta$. Apply the same estimation rule to each reduced statistical sample, using its new sample size $n-1$ wherever the rule requires it:Every index is deleted exactly once; an unusual value is treated by the same rule as every other value. The mean ${\overline{\theta }}_J$ summarizes the deletion estimates. They overlap in $n-2$ observations, so they are generally not independent random variables representing fresh repetitions of the original experiment. Cosma Shalizi's ^[ref] explain this construction and its scaling.

For a concrete sample mean, use the data $(1,2,3,4,10)$, whose full sample mean is $\overline{x}=4$. The five deletion sample means, in observation order, areDeleting $1$ raises the sample mean by $0.75$, while deleting $10$ lowers the sample mean by $1.5$. Thus $10$ has the largest deletion effect in this statistical sample. This is a descriptive sensitivity calculation, not a rule for removing that observation: the full sample mean remains $4$.

Early literature includes M. H. Quenouille's ^[ref] and John W. Tukey's one-paragraph abstract ^[ref], within the issue's collected abstracts. Shalizi's ^[ref] discuss the later development of jackknife bias correction and the jackknife variance estimator.