Assume constant mass density, a prismatic channel, approximately uniform cross-sectional velocity , a hydrostatic pressure distribution, small bed slope , and no lateral inflow. Integrating mass conservation over a slice of length gives , hence .
Let be the hydrostatic first moment, where is the actual free-surface depth. The downstream momentum balance is
Using reduces its left side to . For a fixed prismatic cross-section, , so . Therefore the exact depth-averaged result is
Here hydraulic radius converts perimeter shear stress into force per unit fluid volume. The printed notation yields the requested equation if it denotes this hydraulic pressure head, whose derivative equals .
The geometrical definition printed in the next part needs care. Triangular-channel hydrostatic pressure head gives the factor explicitly. For the stated symmetric triangular notch, the actual depth is , whereas its printed is . Indeed and , giving . Thus the equation with precisely the printed geometrical has pressure term . A first-principles derivation cannot remove that factor. The subsequent intended scaling can be carried out algebraically using the printed convention, as below, or with the pressure scale doubled for the actual triangular channel.
A coaxial shell of radius and thickness has mass . Integrating squared radius against mass, the moment of inertia of a uniform solid cylinder about its axis is
On the smooth incline, no tangential contact force produces spin, so the initially stationary cylinder simply slides with acceleration . Therefore .
For rolling without slipping, let be uphill static friction and the downward translational acceleration. Newton's second law, the rotational torque equation, and the contact constraint give
With , these give and . The rolling-time factor on a straight incline is consequently
The required friction is ; the stated no-slip motion presupposes sufficient static friction. At an instantaneously stationary contact point it does no work. Gravitational potential energy is shared between translational and rotational kinetic energy, explaining why rolling accelerates more slowly than the smooth slide.
For an attractive central force, the transverse acceleration vanishes. In polar coordinates,
Thus the specific angular momentum is constant. Assume , so angle can parameterize the orbit, and put . Primes denote angle derivatives. The chain rule gives
The outward radial component of the attractive force is . Its Newton's second law equation is therefore , giving the Binet equation
The sign reflects being the positive inward force magnitude. Zero angular momentum produces radial motion and is outside this angular parametrization.
Put . In the Unscaled Papkovich–Neuber representation, choose the componentwise harmonic vector field
Both and are harmonic away from the origin: the latter is a linear combination of derivatives of the harmonic function . Also , so the representation gives . The no-slip boundary condition requires and , giving
Thus the Rotational Stokes flow between concentric spheres is
The divergence of is zero: the cross-product field itself has zero divergence, and is radial and perpendicular to . Hence in this pressure gauge.
There is also a symmetry argument. Take the rotation axis as the polar axis. Linearity of Stokes flow makes velocity and pressure perturbations change sign when the rotation reverses. Reflection in any plane containing that axis reverses the axial rotation vector. Combined with axial symmetry, this excludes meridional velocity components and forces the pressure perturbation to be both unchanged and sign-reversed by that reflection. Equivalently a purely azimuthal axisymmetric Stokes flow has no meridional component of its vector Laplacian, so its meridional pressure gradient vanishes. Thus the pressure is constant; setting that arbitrary constant to zero gives the same result without solving the radial coefficients.
For the Newtonian fluid stress tensor, let . The symmetric part of the velocity gradient is
because the gradient of rigid rotation is antisymmetric. Consequently the full stress tensor, with , is
For , its only nonzero spherical-coordinate components are . On the inner sphere the traction exerted by the fluid, using the sphere's outward normal , is
Its torque is
using . Therefore the applied Torque in rotational Stokes flow between concentric spheres is
For , the relevant speed and length are and , so negligible inertia requires the rotational Reynolds number
For a narrow gap , the viscous gradients are controlled by , not . A centrifugal acceleration of scale is then compared with viscous scale , rather than . Their ratio is . This explains why the large-gap condition is not a necessary narrow-gap criterion; a thin gap greatly strengthens viscous resistance. This base-flow scaling is distinct from a criterion for stability to arbitrary short-scale disturbances.
The Minimum-dissipation theorem for Stokes flow compares a Stokes solution with all sufficiently regular incompressible trial velocities in the same fluid domain that have the same prescribed boundary velocities. With no body-force work, the dissipation functional is
If and on the prescribed boundary, integration by parts with and makes the cross term vanish. Thus
The comparison fixes velocities, rather than applied torques or forces.
Now let the particles take their actual rigid translations and rotations in the particle-containing flow. Extend the fluid velocity through each particle by its rigid velocity. No-slip boundary conditions make this extension continuous; rigid motion has zero rate-of-strain tensor and zero divergence. The extension is therefore an admissible trial field in the original particle-free annulus, and its dissipation equals that of the fluid outside the particles. Hence .
The Stokes boundary-work identity gives . The outer sphere is stationary, the maintained inner sphere has zero translational velocity, and every added particle has zero total force and torque, so none of those other boundaries contributes power. Likewise . Therefore
For nonzero rotation and a particle of positive volume the inequality is strict: equality in the dissipation identity would force the extension to be the unique original flow, but that original flow has nonzero strain on every open particle region, whereas a rigid interior has zero strain. This proves that force-free inclusions increase rotational resistance:
No dilute-suspension approximation or assumption about the particle shapes was used.
In a slender symmetric viscous sheet, the two broad faces have zero tangential traction. A cross-thickness variation of on the thickness scale would produce leading shear , whereas is smaller by the square of the aspect ratio. The leading transverse momentum balance and the shear-free conditions therefore give : write .
Incompressibility gives . Symmetry about sets the integration constant to zero, so
The normal traction on either nearly vertical face matches the ambient hydrostatic pressure, giving to leading order. The Newtonian fluid stress tensor then yields
The factor four is the planar extensional resistance, as in the planar viscous-sheet stretching equations, rather than the factor three for uniaxial extension with two contracting transverse directions.
The following slice diagram shows all axial-force contributions per unit -width; the end arrows indicate positive tensile stress and reverse if that signed stress is negative.
Figure 1.
Signed end tractions, ambient side pressure and weight on a widening viscous-sheet slice
. Details.
The two end tractions have net -component . The normal pressure on the inclined side faces contributes , and gravitational acceleration gives the weight . Hence force balance is
Substitute the stress expression and :
so
Keeping the side-pressure component is essential: dropping it would retain an incorrect dependence on the arbitrary ambient pressure level. The remaining equation from conservation of mass is
It follows either by integrating incompressibility across the moving faces or directly from the face kinematic boundary condition.
When effective gravitational acceleration vanishes, is independent of . It equals the applied excess tensile force , with ambient pressure removed from the end loading. Along a material element labeled by its initial position ,
Integration gives uniform material thinning under planar-sheet tension:
A material interval conserves its area: . Thus
With the specified fixed material origin, integrate the latter relation to obtain
These formulas hold while the material thickness remains positive.
For the quadratic initial profile, put . If , evaluating the integral at gives
For it instead gives
For a constant positive pulling force, first reaches the minimum initial thickness at
Both expressions for diverge there. Write . If , the inverse tangent tends to and , giving
For ,
This is finite-time extension of a quadratically thickened sheet. With nonzero , the dominant extension comes from the small material region near the thickness minimum, where the integrand is proportional to . Its width shrinks like , giving an integrated divergence . The fixed ends are asymptotically far away in those local coordinates, so does not depend on . For a uniform sheet the entire material interval has denominator , giving the stronger divergence and retaining its length in . The singularity is a prediction of the ideal slender-sheet model as its minimum thickness tends to zero.
Write , , and use the usual reduced rigid-recoil representation , with zero initial recoil and a smooth stroke returning and to zero. For this representation,
Define the fluid impulse and its first moment by
The force law from part (i) gives, by integration by parts,
These are the endpoint momentum flux in elongated-body recoil. Put
The endpoint quantities are needed in addition to the integrals unless further end-shape assumptions are made. Lateral force and yaw torque balance are
Expanding gives the requested simultaneous second-order equations, with the retained endpoint fluxes:
Substituting and integrating once from the initially straight, unrecoiling state yields
These are first-order coupled equations for with known forcing.
The claimed mass-independent final angle requires a qualification. Let
After the stroke, and . If the post-stroke dynamics are stable, the first-order system relaxes to
Let . Solving this two-by-two system gives
This is quadratic-bend turning with finite body mass. At the limiting state , so . The forward velocity is in the chosen direction, and its lateral component has precisely the body's slope: the fish glides parallel to itself. The angle is generally reached after a relaxation transient, not necessarily immediately at .
The advertised result is obtained in the thin-body approximation that neglects relative to fluid added mass, or in the special geometry :
For example, take negligible nose added mass and nonzero tail added mass , so . In the zero-body-mass approximation, the post-stroke equations for are homogeneous. The inertia matrix is positive definite by Cauchy-Schwarz inequality, and
The two eigenvalues of therefore have positive real parts: their sum and product are positive, whether they are real or conjugate. For the transient decays, proving parallel gliding and the displayed turn in this approximation.
With finite , the angle is not a consequence of the printed force law for arbitrary . A concrete stable counterexample uses , , and . Then , , , , , and . Hence
which differs from for positive body mass. This example has positive determinant and positive matrix trace of the relaxation matrix. Another immediate check is that if both endpoint added masses vanish, integrating the force law conserves total lateral momentum. A finite-mass fish cannot then end with and a nonzero new transverse velocity. Thus endpoint and inertia assumptions cannot silently be omitted.
If is literally the centre of mass position of a deforming body, use a mass-centered deformation rather than the raw quadratic. For body mass density , its mean is , with ; in the slender approximation . The reference translation above is then . Likewise the active angular momentum contains , where , which vanishes for symmetric mass distribution. These corrections modify derivative forcing during the stroke. Since the shape and its time derivative return to zero, they do not remove the finite-body-mass qualification of the final impulse relation.
The principal physical limitations are small slopes and small total turning angle, prescribed approximately constant forward speed, slowly varying slender geometry, and neglected viscosity, separation and detailed three-dimensional wake interactions. Fast large-amplitude C-starts need more than this linear theory. Neglecting body mass is most reasonable for a laterally thin body whose crossflow added mass greatly exceeds its displaced mass. Stable relaxation also requires an appropriate tail/end geometry; it is not guaranteed for every added-mass profile.
The Lighthill elongated-body theory is a reactive, nearly inviscid flow description of a long slender swimmer, appropriate when the longitudinal Reynolds number is large but its lateral displacement and slope are small. To leading order, crossflow past each section is the two-dimensional potential flow generated by lateral motion of that section. Longitudinal variation is slow on the cross-sectional scale, and axial speed is prescribed at this order. With increasing from nose to tail, fluid passes sections in increasing , and the linearized relative lateral velocity is
The section's added mass per unit length is defined by its crossflow kinetic energy . Equivalently, if is the velocity potential for unit lateral section motion, then over the exterior crossflow plane. It has units of mass per length and depends on cross-sectional shape and density. For a circular section of radius , gives . It is not the body's mass per unit length.
The transverse fluid impulse attached to a section is . A control segment has both changing impulse and an impulse flux through its ends. Taking their sum per unit length gives the reaction on the fluid:
The force on the fish is . In particular, is inside the convective derivative; dropping its spatial derivative or the endpoint fluxes is not justified merely because the fish is slender.
The recoil correction in elongated-body theory adds unknown whole-body translation and yaw to a prescribed active bending shape:
They are determined by total lateral linear momentum and yaw angular momentum balance, rather than by imposing that the deforming swimmer remains on its original axis. Body inertia and hydrodynamic added inertia have the same sign. If the translation variable is required to be the literal centre of mass, the active deformation must first have its mass-weighted mean removed: the reference displacement is otherwise not the centre of mass displacement during the stroke. The corresponding deformation contribution to angular momentum must also be retained when it is significant.
Choose an explicit orientation convention, since the supplied PDF does not display the referenced diagram. Let point upward and complete a right-handed frame with the horizontal flow direction . Use
where points from the head towards the tail. A tail point has position . Its rotational velocity is , where the scalar is about and therefore . Keeping that sign is essential for stability.
The point lies at height , so the local imposed fluid velocity is . Its velocity relative to that fluid has components
where and . These follow by projection; rigid rotation has no tangential component.
Write , , , , and
Let . The resistive-force theory force per length on the tail is . Head drag and excess weight give
The fluid vorticity is , so its local angular velocity has scalar component about our axis. Sphere rotation resistance therefore contributes . Torque balance about , including gravity acting at , is
Eliminating gives the unapproximated local-resistance relation
Now retain head and tail translation resistance at the same order, as intended by , but use in the lever-arm integrals. Thus , . Sphere rotation resistance is smaller than tail rotation resistance by order , so omit at fixed scaled shear. Also neglect relative to the tail's hydrodynamic force-centre distance . Uniform density gives
hence this step requires , or head-dominated volume. It is consistent with the distinguished logarithmic ordering, but by itself would not ensure it. The model further neglects hydrodynamic interaction between head and tail, and uses low-Reynolds-number local drag.
Under these approximations,
so
Because , its equilibria satisfy . Define
with at zero shear. The two orientations modulo are and . The derivative of at an equilibrium is
The second factor is positive, so the branch is stable and the other is unstable. On the stable branch the tail tip is above : the head is lower than the far tail end.
The corresponding leading translation velocities are
The second formula uses in the normal force balance. In the equilibrium orientation , and simplifies to
This is shear-dependent settling of a sphere-and-tail body. Since for a slender filament, increases with the magnitude of shear and the signed upward-coordinate velocity increases towards zero. Equivalently, increasing shear decreases the downward sedimentation speed. The formula remains negative throughout this leading model. If there is no shear dependence; reversing the drag anisotropy would reverse the trend. At very large shear, the omitted sphere rotation term proportional to can become important, so the approximation is not uniform in arbitrarily large .
The cell conservation equation has the form , with
The first term is advection by the suspension's fluid velocity, the second is active swimming at fixed speed in direction , and the last is an isotropic diffusive flux due to random swimming. Here has units length squared per time, is cell number per volume, and the divergence measures local loss by transport. No birth or death term is included.
For a spherical cell of radius , its excess buoyancy force is . The Stokes drag law gives passive settling speed . Neglecting individual settling requires , not exact neutral buoyancy. A dilute suspension can still generate appreciable collective buoyancy because many small excess masses contribute to the bulk density.
Use the half-rate convention for the gyrotactic reorientation time , required by the coefficients printed in the question:
The first torque tends to point the bottom-heavy cell upward; the second rotates a sphere with half the fluid vorticity. This is a quasistatic torque balance, assuming orientation relaxation is fast enough to follow the local flow. Keeping orientation dynamics instead would put on the left. The convention with aligning rate uses a reorientation parameter twice this . Thus the factor of two is a definition, not a physical discrepancy. Close to the upward state the quasistatic gyrotactic balance with half-rate reorientation gives .
For a dilute Newtonian suspension with the Boussinesq approximation, the other equations are
The background-water hydrostatic pressure is absorbed in . The cell volume is , density is , cell density is , dynamic viscosity is , and is the volume fraction. Extra cell mass produces the downward buoyancy term. Constant viscosity and no explicit swimming stress are part of this closure. Rigid impermeable plates impose no-slip boundary condition and zero normal total cell flux.
At rest, the stable cell direction is upward and the steady vertical flux is . It is constant in depth by conservation and zero at an impermeable plate, so
The pressure gradient balances its excess weight. The mean-concentration formula follows by averaging over and is continuous at .
Use , , , , and . Then
where
The dimensionless base concentration is .
Linearize with the horizontal Fourier mode and . Write the first-order orientation perturbation as . Unit length gives , while torque balance gives . Incompressibility implies
Let . The perturbation of fluid advection is , and swimming contributes . Cell conservation therefore gives
or
This explains the destabilizing concentration-advection term and the gyrotactic redistribution term with their signs.
To eliminate pressure from the fluid equations, take their divergence, giving . Apply to the vertical equation . The concentration second derivatives cancel, leaving
No-slip conditions give and at each plate; for nonzero , the latter follows from the vanishing horizontal velocity and incompressibility. Zero perturbed normal cell flux gives
The normal orientation correction is zero at this order, and the fluid velocity is zero at the plate, so neither adds another term to this concentration condition.
Now use the stated distinguished scalings , , , , , and . Here is a scaled wavenumber, unrelated to Question 2's shear rate. At order one, the concentration equation and boundary conditions give , at both endpoints, so is constant. Absorb its nonzero value into the perturbation amplitude and choose
At order , the fluid equation is with at both ends. Its unique solution is
A homogeneous difference is cubic and the four clamped conditions make it zero.
The concentration equation at order gives
so , where fixes the amplitude normalization. At order it gives
The boundary conditions expand to at each endpoint. Integrating this equation from zero to one cancels against . Also and
Thus the required solvability condition is
Since , the clamped-quartic solvability in shallow bioconvection gives
For nonzero long horizontal waves, gives decay and gives growth; the leading neutral threshold is . The leading instability is stationary and independent of and . The exactly uniform horizontal mode has zero growth at this order because total cell number is conserved, so it does not determine the threshold. Upward swimming produces a dense upper layer; the unstable disturbance is its buoyancy-driven overturning, or bioconvection. Higher orders are needed for wavelength selection and finite-depth threshold corrections.
With fixed material parameters and mean concentration, for small , so a sufficiently thin chamber is stable to this branch. Formally its threshold corresponds to when the weak-stratification and distinguished-gyrotaxis assumptions apply. In particular, the assumed is a specified asymptotic analysis; at fixed and decreasing , one instead has . The leading result should not be asserted unchanged in that different ordering without redoing the expansion.
Apply the reciprocal identity with flow1 the actual body-force-driven flow and flow2 the rotlet auxiliary field. Its body force is zero. The large-sphere boundary terms vanish under the usual decay and convergence assumptions for this unbounded body-force problem.
On the sphere, the actual rigid velocity is . The auxiliary force is zero and its torque with the outward fluid normal is . The actual torque with that normal is the negative of the hydrodynamic torque on the sphere, hence zero for a torque-free sphere. Consequently the surface side of the reciprocal identity is just . The volume side is
Since the auxiliary couple is arbitrary,
An unknown translation contributes nothing because the auxiliary rotlet has zero net force. This is body-force-driven rotation of a torque-free sphere.
Let . The first sphere's leading velocity field is . Taking its curl gives
The second sphere is torque-free, so
Its own force-free and torque-free motion emits no Stokeslet or rotlet; further reflections are smaller at large separation. In particular it spins in the opposite direction for a primary rotation perpendicular to the line of centers, and in the same direction with twice that magnitude for a parallel rotation.
Now isolate the additional interaction caused by the applied force . Its leading Stokeslet produces at the first center the incident velocity . The Stokes drag law makes the increment of the force required to hold that sphere fixed
The sign opposes the incident velocity. This holding force acts on the fluid and generates a reflected Stokeslet. A Stokeslet of force has vorticity . Evaluating half that vorticity at gives
Thus its magnitude is , as required. This is forced-sphere rotation reflected from a held sphere. Longitudinal forcing has zero leading spin by symmetry. Finite-size multipoles and the couple correction needed to preserve the first sphere's prescribed spin are smaller than this force-monopole reflection by order .
Measure pressure relative to its hydrostatic value and let be the lower-side excess pressure minus the upper-side excess pressure. The long unobstructed parts of the tube act as Poiseuille resistances in series. Their lengths add to at leading order because ; entrance and exterior resistances are smaller for . Consequently
Here is downward laboratory-frame volume flux.
Take upwards in the particle frame and let measure distance from the particle surface to the tube wall. In this frame the particle is stationary and the wall moves upwards at speed . In the thin, slowly varying gap, pressure is uniform across and the leading axial Stokes equation is . No slip gives
The gap's upward relative flux is constant, while conservation of displaced volume gives . Thus , and integration from bottom to top yields
Curvature corrections to the gap circumference are relative order . Rapid end regions do not contribute to the leading singular neck resistance.
For the force use a cylindrical fluid control volume containing the particle, with full-area end caps just beyond it and the particle surface as an inner boundary. The upward pressure force on the caps is . The outer wall exerts axial traction on the fluid, whereas the particle's traction on the fluid integrates to minus its upward hydrodynamic resistance . The zero-inertia momentum balance is therefore
The velocity profile gives , proving
Using the outer-wall traction is essential; directly using the particle-side shear and a flat-end pressure force would omit the pressure on the sloping particle surface. This is control-volume drag formula for a sphere in a tube.
The four dimensionless relations are
Solving them gives
Within the preceding lubrication and tube-resistance approximations, the force is
Since , . Dropping the smaller additive terms in the numerator yields the requested leading formula
The Cauchy-Schwarz inequality gives , so the retained term is positive. The term multiplied by must be retained: it can compete with even though the unmultiplied lower integrals cannot.
For a sphere of radius , the neck has at leading order. With , the integrals reduce to . For these last integrals are , giving the full two-sided integrals. Put
The printed sphere integrals are half these values: they correspond to integrating over one side of the neck rather than the stated interval spanning both sides. The substitution above establishes the radius-deficit normalization of sphere-tube lubrication integrals. Using the two-sided values, the force becomes
so the three answers are
These are the corrected sphere-tube drag regimes with a radius-deficit gap. If the supplied half-integrals are treated as stipulated data instead, the same algebra gives , and in the three regimes. Thus the first and third numerical drag coefficients double when the geometrically correct integrals are used; the regime boundaries and dominant flow patterns are unchanged.
In the first regime, , , and . Almost no displaced fluid escapes through the long tube; it bypasses upwards through the thin annulus. The large pressure-driven bypass resistance sets the drag, with shear smaller.
In the second regime, , , and
Almost all displaced volume travels through the unobstructed tube, whose pressure resistance dominates the force. Both terms in this small gap flux are worth displaying: their crossover at changes the balance between pressure-driven and wall-driven gap transport but does not create a fourth leading force regime.
In the third regime, , , and . Tube throughflow still carries almost all the displaced volume, but the dominant resistance is local annular lubrication rather than the net tube pressure drop. The relative gap flux is now of order , so wall-driven and pressure-driven components both matter locally. The formulas describe hydrodynamic resistance after subtracting hydrostatic buoyancy; terminal settling would balance it against the particle's excess weight.
Use the material-first nominal stress tensor convention , whose reference-volume power is . At a regular constraint point, , all admissible deformation rates satisfy . Equality of mechanical power and the derivative of the strain energy density gives
for every vector in this eight-dimensional tangent hyperplane. Its orthogonal complement is the span of . Consequently the constraint reaction in hyperelastic stress is
The scalar Lagrange multiplier is not determined by the work identity: its contribution vanishes on every allowed rate. Regularity matters. For example, replacing a regular incompressibility constraint by would give zero gradient on the same admissible set and could not represent an arbitrary pressure reaction through that gradient.
For incompressibility use the regular function , with . Differentiating the alternating-tensor expression for gives
The first expression is the cofactor entry; contracting it with gives , proving the inverse formula. On this yields
With this plus-sign convention is the negative of the usual pressure multiplier.
For an equibiaxially stretched sheet, incompressibility requires the principal stretches . The printed sheet expression's is inconsistent with this constraint and must be ; the balloon expression later on the same PDF page has the correct exponent. Let denote the derivative with respect to principal stretch , evaluated on this constrained path. With no applied thickness-direction traction,
Isotropy gives at equal in-plane stretches. Hence
The equibiaxial nominal tension of an incompressible sheet is therefore
The two equal in-plane nominal tensions do work per reference volume; the thickness-direction nominal stress does no work because it is zero. That is exactly the constrained energy differential, explaining energy conservation. As a check that the printed exponent is a genuine error, a neo-Hookean solid gives , zero at the undeformed state. The derivative with the erroneous exponent would instead give at that state.
For the thin spherical balloon, use the leading reference shell volume . Its two tangential stretches are , while its thickness stretch is and its current thickness is . Define . Its total elastic energy at leading thin-shell order is
During a quasistatic increment , the enclosed volume changes by , and . Equating energy change to work of the internal excess pressure gives
This is the inflation pressure of a thin incompressible elastic balloon. The approximation neglects through-thickness variation and curvature corrections beyond leading order, and requires . It also agrees with membrane force balance: , , and . Here pressure is force per current area, while is force per reference area; confusing them would lose a factor of .
Approximate the gap by steady Couette flow, neglecting edge effects and additional losses from membrane deformation. Its velocity gradient is , so the shear stress is . The force on the gliding bacterium and the rate of viscous dissipation are
The same power follows by integrating over gap volume . In SI units, , , and . Thus
An energy needs a duration or distance: for in seconds, equivalently for travel distance . Moving one micrometre takes one second and dissipates . This is the mechanical cost of the sheared film. A numerical biological efficiency would additionally require the motor's input power and the losses associated with its peristaltic actuation.
The compressive tip force is limited by Euler buckling of an elastic filament. With unsupported length and filament bending modulus ,
For a hollow microtubule, use rather than the solid-cylinder value. For a clamped seed and a moment-free tip free to slide along the wall, linearized angle balance is , with and . Its first nonzero solution has , hence . A different tip constraint changes this coefficient.
Increasing the unsupported length decreases that threshold. For growth against a wall at fixed projected gap , polymerization-driven cantilever postbuckling gives a more direct quantitative statement. The inextensible elastica obeys . Multiplying by and using the zero tip moment gives
Put and substitute . Integrating arc length and its horizontal projection gives
where the complete elliptic integrals are and . Using and gives , and therefore
Thus the initial postbuckling force decreases as more contour is inserted. This result describes the weakly buckled branch before overhang, further wall contact, or altered tip constraints invalidate the model. It is a mechanical force law; a chemical force limit from polymerization can impose a smaller force.
Set , , and . To keep dimensions and the thermal amplitude explicit, write the force covariance as . The printed unit coefficient is a choice of noise normalization, not a general dimensional thermal force. Assume the initial velocity is independent of the future white noise. The Underdamped Langevin dynamics has integrating-factor solution
Taking its expectation and using the delta covariance in the double integral gives
Here the noise contribution is . The equipartition theorem requires the long-time value . Consequently the fluctuation-dissipation relation for a Langevin particle is
With an initially thermal velocity, and , so the velocity variance is stationary. If one insists on the printed coefficient in dimensional units, the model has temperature fixed by in those units; it cannot describe arbitrary temperature while retaining that coefficient.
The numerical contour length is , and . Begin with a three-dimensional freely jointed chain in which each chemical monomer is also an independent statistical link of length . This neglects excluded volume, hydrodynamic effects on equilibrium statistics, and stretching of the chemical backbone.
An applied force gives a one-link canonical partition function
The mean extension is . Expanding the Langevin function at small force gives . The three-dimensional entropic chain stiffness is thus
The unforced root-mean-square end-to-end distance is , not the contour length. This response is entropic straightening of a fluctuating coil. Axial stretching of a straight material rod instead has stiffness , and is a different phenomenon. Aqueous solvent does not by itself prove ideal-chain statistics: a good-solvent self-avoiding walk has a different coil size and susceptibility. The estimate above states explicitly the model under which the supplied link data suffice.
An optical tweezers force requires the change in optical momentum. Define the optical momentum efficiency by , with all ray geometry and medium-index factors absorbed in and the vacuum light speed. The specified relative refractive index does not determine without bead size and the beam's convergence, waist, and ray directions. A unique exact force is therefore not supplied by these data.
Retaining the entropic stiffness from part (a), the parameterized answer is
Using the persistence-length-corrected worm-like chain instead gives in its small-force regime.
For a definite scale estimate, take the weak-contrast optical momentum change to be . This gives , for part (a)'s model, or for the corrected chain. These are estimates with an explicitly chosen ray-momentum efficiency, not a universal formula for a transparent trapped sphere.
Another illustrative optical model makes the uncertainty visible: normal-incidence Fresnel reflectance is . An effective reflector with in the simple momentum convention gives and extensions or , respectively. A refracting bead in a focused beam is not just that reflector; entry and exit rays also transfer momentum. Neither estimate eliminates the missing optical geometry.
The extension must be compared with the actual contour length, . For the illustrative force and the persistence-length-corrected chain, and . Both are small, so the small-force Langevin function expansion and entropic-extension approximation are consistent. Although the mean extension greatly exceeds the unforced millimetre-scale coil size, it remains far below the maximum contour length; that does not violate the model.
The material axial elongation at this force would be only , illustrating that most of the predicted coil extension is orientation and straightening rather than stretching the backbone. Thus the order-of-magnitude entropic response is internally reasonable within the ideal-chain assumptions, but its numerical optical-force prediction is not unique. Solvent-dependent excluded volume, the unusually large stated contour length, actual trap geometry, and surface interactions all matter to an experimental interpretation.
The viscosity converts to , and the translation speed is . Mechanical power is . The hydrodynamic drag models for a polymer show why a conformation assumption is necessary.
For a largely aligned slender filament translating along its axis in unbounded fluid, use
With , , so . The required force is about , and
Transverse translation has roughly twice the leading logarithmic drag. This estimate treats the entire contour as an aligned slender body, assumes creeping flow, and omits bead drag, wall corrections, and additional motor losses. It is not a drag formula for an unstretched coil.
For an alternative nondraining coil, approximate the hydrodynamic radius by its radius of gyration, . The Stokes drag law gives and . A freely draining toy model that sums spherical-monomer drags of radius instead gives . Hydrodynamic interactions are suppressed in this last model. These distinct answers explain the need to specify conformation and hydrodynamic coupling; the stated data do not select a unique one.